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  • Reference article Jørgensen, P. M. & S. León-Yánez. (eds.) 1999. Catalogue of the Vascular Plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: i–viii, 1–1181.
  • Reference article Lawesson, J. E., H. Adsersen & P. Bentley. 1987. An updated and annotated check list of the vascular plants of the Galapagos Islands. Rep. Bot. Inst. Univ. Aarhus 16: 1–74.
  • Reference article Porter, D. M. 1983. Vascular plants of the Galapagos: Origins and dispersal. 33–54. In M. B. R. I. Bowman & A. E. Leviton Patt. Evol. Galapagos Org.. Pacific Division, AAAS, San Francisco.
  • Reference article Wiggins, I. L. & D. M. Porter. 1971. Fl. Galápagos Isl. 1–998. Stanford University Press, Stanford.
 
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