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(Last Modified On 8/7/2013)
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(Last Modified On 8/7/2013)
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Species
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Gasteranthus acropodus (J. D. Smith) Wiehl.
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PlaceOfPublication
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Selbyana 1: 155. 1975.
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Synonym
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Besleria acropoda J. D. Smith, Bot. Gaz. (Crawfordsville) 54: 240. 1912. TYPE: Forests of Tsaki, Talamanca, Prov. Limon, Costa Rica, Tonduz 9554 (US, holotype). Besleria allenii Morton, Ann. Missouri Bot. Gard. 29: 38. 1942. TYPE: Panamai, Allen 2371 (US, holotype; MO, isotype). Besleria allenii var. paucivenia Morton, Ann. Missouri Bot. Gard. 29: 38. 1942. TYPE: Panama, Allen 2188 (US, holotype; US, isotype). Gasteranthus allenii (Morton) Wiehl., Selbyana 1: 155. 1975.
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Description
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Terrestrial subshrubs or shrubs; stems erect, to 1(-5) m tall, the younger stems strigillose, the older stems glabrescent. Leaves subequal in a pair, more or less elliptic, 7-20 cm long, 3-7 cm wide, membranous, the apex acuminate, the base obtuse to attenuate and decurrent, serrulate or crenulate to remotely serrate, above dark green and glabrous, below pale green and sparsely strigillose; petiole 1-3 cm long, strigillose. Inflorescences axillary but appearing terminal by re- duction of the main axis, umbellate or cymose, few flowered; peduncles 1.8-5.5 cm long, sparsely strigillose; pedicels 0.5-2.0 cm long, sparsely strigillose. Flowers with a broad receptacle, the calyx zygomorphic, the sepals distinct, un- equal, serrate to nearly entire, whitish, pale green or purplish, the posterior lobe ovate or rhombic, keeled, 9-15 mm long, the 2 lateral lobes obliquely ovate, 8-15 mm long, the 2 anterior lobes lanceolate, slightly keeled, 6-15 mm long; corolla oblique in the calyx, mainly yellow, but varying from cream white to lemon yellow, with or without reddish lines, spots or patches, the tube ventricose and broad from a spurred base, 2.2-2.8 cm long, the spur 6-8 mm long, the tube to 1.5 cm broad at the middle, slightly narrowed just below the limb, sparsely pilose outside the limb, slightly bilabiate, ca. 1.5 cm broad, the lobes ca. 2-5 mm long; stamens 4, included, the staminode short, attached to the corolla base below the nectary, the filaments flattened, curved, the anthers 1-2 mm long, con- nate; disc thickened posteriorly, pubescent; ovary glabrous to sparsely pilose, style ca. 7 mm long, pubescent, stigma bilobed. Fruit oblate globose, ca. 5 mm long, 8 mm wide; seeds oblong, light brown, obliquely striate.
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Habit
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subshrubs or shrubs
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Note
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Gasteranthus allenii had been considered distinct from G. acropodus Wiehl because it lacked the spots on the corolla. However, from examination of recently collected material over the distributional range, it has become evident that corolla coloration is quite variable.
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Distribution
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known from both Costa Rica and Panama, inhabiting moist forests mainly on the Caribbean slope.
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Specimen
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BOCAS DEL TORO: Hill above RR station at Milla 7.5, Croat & Poiter 16378 (MO). Hill- side above Almirante, Gentry 2746 (MO). Quebrada Huron, Kirkbride & Duke 432 (MO). Chiriqui Trail between Criollo, just above Buena Vista & Quebrada Higueron, Kirkbride & Duke 791 (MO). Water Valley, Wedel 1420 (MO, US), 1475 (MO, US). Fish Creek Mountains, Wedel 2272 (MO), 2346, 2347 (both MO, US), 2351 (MO). Fish Creek Hills, Wedel 2455 (MO, MO). COCLE': N rim, El Valle de Anton, 600 rn, Allen 1829 (MO, US). Hills N of El Valle de Anton, 1000 m, Allen 2188 (US, US). La Mesa, N of El Valle de Anton, 1000 m, Allen 2371 (MO, US). La Mesa above El Valle, 810 m, Croat 25304 (MO). Hills N of El Valle de Anton, Dressler 4388 (PMA). Cerro Gaital Caracoral, near Cerro Pilon, 2700-3200 ft, Dwyer & Correa 8851 (MO, US). Mountains N of El Valle de Anton, 2500-3000 ft, Lewis et al. 1755 (DUKE, MO, SCZ). COLON: Rio Guanche 3-5 mi inland, 10-200 m, Croat 26197 (MO), 26209 (PMA), 50-200 m, Croat 36990 (MO); Folsom 3767 (MO, US); Foster et al. 2339 (DUKE); Mori et al. 6441 (MO); Mori & Kallunki 3012, 3123 (both MO). Along old logging road above Rio Iguanita, dense second growth forest, 390 m, Skog et al. 4185 (US). PANAMA: Cerro Campana, Croat 17231 (US); Dressler 2984 (PMA), 3520 (PMA); Kirkbride 240 (MO, NY); Mori & Bolten 7684 (MO). SAN BLAS: SE of Puerto Obaldia, Croat 16771 (MO).
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