2. Carya cordiformis (Wangenh.) K. Koch (bitternut hickory, pignut hickory)
Pl. 430 g, h;
Map 1924
Plants trees to
35 m tall (to 50 m elsewhere). Bark gray to brownish gray, smooth or shallowly
furrowed and exfoliating in small flakes. Twigs 3–4 mm thick, tan to light
brown, the terminal bud 4–11 mm long, yellow or tan, the bud scales meeting at
their margins and not or scarcely overlapping, the bractlets around the
axillary buds free (but usually completely hidden by dense peltate scales).
Leaves with the petiole and rachis pubescent (sometimes becoming nearly
glabrous toward the petiole base), with (5)7 or 9 leaflets. Leaflets 4–18 cm
long, 1–8 cm wide, lanceolate to oblanceolate or obovate, straight or weakly
arched (appearing slightly asymmetrically tapered), the margins finely to
coarsely toothed, glabrous or with scattered hairs, the upper surface glabrous
or with scattered, small, circular, whitish to brownish, peltate scales, the
undersurface with scattered to relatively dense, branched (the branches
appearing fasciculate) hairs, sometimes mainly along the veins or becoming
nearly glabrous with age, and also with scattered, small, circular, pale yellow
to brownish, peltate scales. Staminate catkins noticeably stalked to nearly
sessile. Fruits 2–3 cm long, 2–3 cm wide, globose to somewhat ellipsoid, not
flattened, with low wings along the 4 sutures, the husk 1–2 mm thick, splitting
50–80% of its length, with small golden yellow scales that wear off with age.
Nut not or only slightly flattened, the shell less than 1 mm thick. Seed
bitter. 2n=32. April–May.
Scattered to
common nearly throughout the state (eastern U.S. west to Minnesota and Texas;
Canada). Banks of streams and rivers, margins of ponds, lakes, sinkhole ponds,
oxbows, and sloughs, bottomland forests, bases of bluffs, and less commonly
mesic upland forests; also roadsides.
Plants with
unusually broad leaflets (5 cm or broader) have been called f. latifolia
(Sarg.) Steyerm.