1. Lamium amplexicaule L. (henbit)
Pl. 434 f, g;
Map 1950
Stems 4–35 cm
long, erect or ascending, often from a spreading base, unbranched or
few-branched from at or near the base, glabrous or more commonly sparsely to
moderately pubescent with short, downward-bent hairs. Leaves short- to
long-petiolate (lower and often also median leaves), the shorter petioles often
narrowly winged, the longer petioles usually unwinged, grading into sessile
leaves (mostly those subtending flower clusters) that sometimes appear more or
less clasping the stem. Leaf blades 5–15 mm long, unlobed or 3-lobed, the
margins also with relatively coarse, blunt teeth or scallops, rounded to
shallowly cordate at the base, rounded to bluntly pointed at the tip, the
surfaces variously densely soft-hairy to nearly glabrous, those of the lower
leaves ovate-triangular to broadly ovate or nearly circular, grading into those
of the upper leaves, which are nearly semicircular to very broadly ovate or
kidney-shaped, usually wider than long. Inflorescences with most of the nodes
well-separated, the bractlets absent. Calyces 5–7 mm long, the tube and lobes
densely bristly-hairy, the lobes shorter than to slightly longer than the tube.
Corollas (except in cleistogamous flowers) 10–20 mm long, the outer surface
sparsely to more commonly densely hairy, with a very dense patch of usually
longer, darker hairs on the upper lip, the upper lip 3–5 mm long, rounded to
shallowly notched at the tip, pinkish purple, rarely entirely white, the lower
lip 1.5–3.0 mm long, usually lighter pink or white, with purple spots or mottling,
rarely entirely white, the lateral lobes reduced to small convexities along the
lip margin, the central lobe obcordate. 2n=18. February–May, rarely
November–January.
Scattered to
common nearly throughout the state, but apparently less common in most of the
Glaciated Plains Division (native of Europe, Asia, Africa; introduced nearly
throughout temperate North America). Edges of marshes, and openings of mesic
upland forests; also crop fields, fallow fields, pastures, fencerows, lawns,
sidewalks, railroads, roadsides, and open, disturbed areas.
Mennema (1989)
treated L. amplexicaule as a complex of five varieties. In his
classification, the introduced Midwestern material would correspond to var. amplexicaule.
However, some of these varieties probably are better treated as separate
species. More recently, based on a molecular-phylogenetic analysis of the
genus, Bendiksby et al. (2011) determined that some of these non-Missouri taxa
previously included within L. amplexicaule represent allopolyploid derivatives
resulting from past hybridization between that species and other members of the
genus. The situation requires further study. Rare plants with white corollas
have been called f. albiflorum Dw. Moore (var. album Pickens and
M.C.W. Pickens) and plants with all of the flowers cleistogamous have been
called var. clandestinum Rchb.
Burrows and Tyrl
(2001) reported that in Australian pastures livestock sometimes contract
neurological problems (staggers) following ingestion of henbit, but that
similar cases of toxicity are unknown from the native range of the species or
from North America.