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  • Reference article Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera. 2012. Rubiaceae a Verbenaceae. 4(2): i–xvi, 1–533. In G. Davidse, M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera (eds.) Fl. Mesoamer.. Missouri Botanical Garden, St. Louis.
  • Reference article Martínez Gordillo, M., B. Y. Bedolla-García, G. Cornejo Tenorio, I. Fragoso-Martínez, M. R. García Peña, J.G. González-Gallegos, S. I. Lara Cabrera & S. Zamudio Ruíz. 2017. Lamiaceae de México. Bot. Sci. 95(4): 780–806. electronic publication
  • Reference article Martínez Salas, E. M., M. Sousa Sánchez & C. H. Ramos Álvarez. 2001. Región de Calakmul, Campeche. Listados Floríst. México 22: 1–55.
  • Reference article Standley, P.C. & L. O. Williams. 1973. Labiatae. En: Standley, P.C. & Steyermark, J.A. (eds.), Flora of Guatemala - Part IX, Number 3. Fieldiana, Bot. 24(9, 3): 237-317. View in Biodiversity Heritage Library
 
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