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(Last Modified On 10/7/2013)
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(Last Modified On 10/7/2013)
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Species
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Heteropteris macrostachya Juss.
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PlaceOfPublication
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Malpigh. Synopsis (Ann. Sci. Nat., Bot. Ser. 2. 13:) 275. 1840.
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Note
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TYPE: Guiana, Herb. Richard (P, not seen; F, photo FM 35589).
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Synonym
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Heteropteris apiculata Miq., Stirp. Surinam Sel. 81. 1851. TYPE: Surinam, Fl. Marowyne, Kappler 1915 (not seen; F, type photo FM 12768). Banisteria apiculata (Miq.) C. B. Robinson ex Small, N. Amer. Fl. 25: 136. 1910.
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Description
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Liana; stems densely pale reddish-brown, pubescent when young, glabres- cent, weakly lenticellate. Leaves opposite, elliptic to lanceolate, gradually to abruptly acuminate at the apex, obtuse to rounded at the base, (4-)8-15 cm long, (2.5-)4-7 cm broad, firmly coriaceous, glabrous and semi-glossy above, pale and tan or golden sericeous beneath, the lower surface sometimes with glands near the base; petioles 5-11 mm long, canaliculate, densely and obscurely appressed- pubescent, bearing 1 or 2 pairs of conspicuous plate-shaped glands on the mar- gins. Inflorescence terminal or upper axillary, paniculate, the branches to 15 cm long, the branches, peduncles, pedicels and calyces reddish-brown, tomentose;
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Habit
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Liana
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Description
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peduncles to ca. 7 mm long, bibracteolate at the apex, the bracteoles pubescent, stiffly erect, to ca. 1 mm long; pedicels less than 2 mm long, articulate to the peduncle. Flowers ca. 1 cm diam., calyx lobes erect, narrowly ovate, 4 of them bearing conspicuous fleshy, gray glands, 2-3 mm long, glabrous on the outside, the thin surface rupturing, at least on drying, densely puberulent within; petals yellow, oblong-elliptic, 3.5-4 mm long, 2.2 mm wide, stamens to 3 mm long, the staminal tube glabrous to sparsely villous, the free part of the filaments less than 1 mm long, the anthers ca. 1 mm long; styles to ca. 2 mm long, glabrous, the apex truncate, triangular, the ovaries ovoid, 2-4 mm long, densely reddish-brown pubescent.
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Note
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According to Johnston (1949) on San Jose Island the species does not produce a typically samaroid fruit. He reported that the wingless ovaries probably act as a fruit. The species is most easily confused with H. obovata with which it shares a thick sericeous indumentum on the lower blade surface. It is distinguished from H. obovata by having thicker leaves with petioles bearing a pair of glands at the middle and by having short pedicels articulated with a bibracteolate peduncle. The pedicels of H. obovata are sessile, not borne on a bibracteolate peduncle. In Panama, the species flowers from October to January but also has been collected in July on Coiba Island.
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Distribution
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ranges from the southern West Indies to northern South America.
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Note
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In Panama the species is known only from tropical moist forest on the Pacific slope.
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Specimen
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DARIEN: Between Rio Punusa and Rio Pucro, Duke 14643 (MO, OS, PMA, US). PANAMA: El Llano, Duke 5845 (MICH, MO). Cerro Azul, 20 km from Panamerican Highway, Dwyer 2208 (MO, US). San Jose Island, Johnston 388, 1039, 1377 (all GH, MO, US). VERAGUAS: Isla Coiba, Campaguja, Dwyer 2369, 2382 (both DUKE, MO). Playa Rosario, northern tip of Coiba Island, Foster 1595 (MO, PMA).
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