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  • Reference article Atwood, J. T. 1989. Orchids of Costa Rica. Part 1. Icon. Pl. Trop. 14: i–iv, t. 1301–1400.
  • Reference article Dodson, C.H. 1992. Checklist of the Orchids of the Western Hemisphere. draft:.
  • Reference article Dressler, R. L. 2003. Orchidaceae. En: Manual de Plantas de Costa Rica. Vol. III. B.E. Hammel, M.H. Grayum, C. Herrera & N. Zamora (eds.). Monogr. Syst. Bot. Missouri Bot. Gard. 93: 1–595. BHL
  • Reference article Funk, V. A., T. H. Hollowell, P. E. Berry, C. L. Kelloff & S. Alexander. 2007. Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contr. U.S. Natl. Herb. 55: 1–584. View in Biodiversity Heritage Library
  • Reference article Hokche, O., P. E. Berry & O. Huber. (eds.) 2008. Nuevo Cat. Fl. Vasc. Venez. 1–859. Fundación Instituto Botánico de Venezuela, Caracas.
  • Reference article Pupulin, F. 2002. Catálogo revisado y anotado de las Orchidaceae de Costa Rica. Lankesteriana 4: 1–88.
  • Reference article Ulloa Ulloa, C., H. M. Hernández, G. Davidse, F. R. Barrie & S. Knapp. 2023. Orchidaceae. 7(2): I-XXI, 1-842. In C. Ulloa Ulloa, H. M. Hernández Macías, G. Davidse, F. R. Barrie & S. Knapp Fl. Mesoamer.. Missouri Botanical Garden, St. Louis.
  • Reference article Ulloa Ulloa, C., P. Acevedo-Rodríguez, S. G. Beck, M. J. Belgrano, R. Bernal, P. E. Berry, L. Brako, M. Celis, G. Davidse, S. R. Gradstein, O. Hokche, B. León, S. León-Yánez, R. E. Magill, D. A. Neill, M. H. Nee, P. H. Raven, H. Stimmel, M. T. Strong, J. L. Villaseñor Ríos, J. L. Zarucchi, F. O. Zuloaga & P. M. Jørgensen. 2017. An integrated assessment of vascular plants species of the Americas. Science 358: 1614–1617 [online suppl. materials: 1-23 + 1-2497], t. 1-4 [f. S1-5].
  • Reference article Ulloa Ulloa, C., P. Acevedo-Rodríguez, S. G. Beck, M. J. Belgrano, R. Bernal, P. E. Berry, L. Brako, M. Celis, G. Davidse, S. R. Gradstein, O. Hokche, B. León, S. León-Yánez, R. E. Magill, D. A. Neill, M. H. Nee, P. H. Raven, H. Stimmel, M. T. Strong, J. L. Villaseñor Ríos, J. L. Zarucchi, F. O. Zuloaga & P. M. Jørgensen. 2018 (onwards). An integrated Assessment of Vascular Plants Species of the Americas (Online Updates).
 
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