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Project Name Data (Last Modified On 9/19/2013)

Flora Data (Last Modified On 9/19/2013)
Species Biophytum dendroides (H.B.K.) DC.
PlaceOfPublication Prodr. 1: 690. 1824.
Synonym Oxalis dendroides H.B.K., Nov. Gen. Sp. P1. 5: 194. 1821. TYPE: Colombia, near Quebrada de la Carbonera, Bonpland 1744 (P, holotype; B, P, isotypes). 0xalis viva Willd. ex Steud., Nom. ed. 2. 2: 242. 1844. TYPE: Colombia, Bonpland 1744 (B, holotype; P, isotypes), nom. illeg. incl. typ. sp. prior. Toddavaddia dendroides (H.B.K.) Kuntze, Rev. Gen. P1. 1: 96. 1891. Biophytum mexicanum Kl. ex Knuth, Pflanzenr. 130: 401. 1930, nom. nud. in syn. Biophytum longiscapum Kl. ex Knuth, Pflanzenr. 130: 401. 1930, nom. nud. in syn. Oxalis dendroides fo. longiscapa Zucc. ex Knuth, Pflanzenr. 130: 140. 1930, nom. nud. in syn. Biophytum mimosoides sensu auct., non (St. Hil.) Guillaumin. Biophytum dormiens sensu auct., non Mart. & Zucc.
Description Rhizomatous herb, stems obsolete or to 35 cm tall and woody, to 5 cm diam.; rooting up to 20 cm from the base, sometimes 2- or 4-branched; bark rough, appressed retrorse pubescent, glabrescent; root fibrous, to 10 cm long, twisted, branched. Leaves pseudoverticillate at the end of the stems, 15-30, 9-26 jugate, linear in outline, subcuneate; lower leaflets ovate to subtriangular, asymmetric, 1-2.5 mm long, 1-2.5 mm wide, the middle leaflets subrectangular to subfalcate, 2.5-10 mm long, 1.5-7 mm wide, the terminal leaflets semiobovate, asymmetric, basally truncate to subhastate, 6-11 mm long, 2.5-5 mm wide, the leaflets some- times overlapping, glabrous or sparsely hirsute on one or both sides, hirsute ciliate, obtuse or subacute, the mucro sometimes violaceous, the midrib medial or emerging from nearer the posterior angle, secondary nerves ascending, curved, branched, connected to a marginal nerve; rachis 2-10 cm long, hirsute, more so at the insertion of the leaflets; petiolules 1/4 mm long, thick, fleshy; petiole 1 mm
Habit herb
Description long, fleshy. Inflorescences subsessile or pedunculate, shortened, glomeruliform or fusiform, only 1 flower appearing at a time; peduncle erect, 1-8 cm long, subappressed pubescent or hirsute, the apex thickened, densely pilose; bracts subulate, to 6 mm long, purplish, pubescent; bracteoles 2-7 mm long, 0.5-1.5 mm wide, triangular linear to linear, the inner bracteoles prominently keeled, the keels somewhat lengthened, ascending ciliate; exceeding or about equalling the pedicels; pedicels 1-5 mm long, articulate in the lower half, glabrous or finely pubescent. Flowers with the sepals linear, 5-7 mm long, 1-1.5 mm wide, acu- minate, 7-10-nerved, glabrous or ascending pubescent, the base densely hirsute, the apex ciliate; petals whitish to pink or lilac, the base obovate or spathulate, 1.5-1.3 times the calyx length, slightly retuse, the base clawed, white; stamens with the filaments basally enlarged, glabrous, the longer stamens 4.5-5.5 mm long, the shorter stamens 3-4 mm long; pistils micro- and macrostylous, the ovary apically pilose, the carpels 1-6-ovuled, the styles sometimes unequal, pilose be- low, the stigma small, bifid. Capsule subglobose or oblong, 2.5-4 mm long, yel- lowish to violaceous, scarious, apically pubescent, or rarely pubescent overall, exceeded 1.75-2 times by the calyx, the carpels 1-6-seeded, short pubescent within; seeds semiovoid-triquetrous, one face more or less plane with an acute margin and the opposite face convex, 1.5 mm long, acute, longitudinally tuber- culate.
Distribution ranges along the American cordillera from Mexico to Ecuador.
Note It has been introduced with greenhouse plants to botanic gardens on other con- tinents, in some cases becoming subspontaneous, for example in Malasia and Brazil. However, records from Knuth (1930) for Brazil are not this species.
Specimen COCLE: Mountain beyond La Pintada, 400-600 m, Hunter & Allen 615 (F, GH, MO, NY, P, U, US).
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