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Published In: Species Plantarum 1: 85. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 8/28/2009)
Acceptance : Accepted
Project Data     (Last Modified On 7/9/2009)
Status: Native

 

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1. Hordeum jubatum L. (squirreltail, foxtail barley)

Pl. 187 c, d; Map 758

Plants perennial, short‑lived, often flowering the first season and thus appearing annual. Flowering stems 30–70 cm long, the nodes dark brownish black. Leaf sheaths glabrous or with spreading hairs. Leaf blades 3–15 cm long, 1.5–4.5 mm wide, roughened or hairy, with a pair of inconspicuous (less than 0.5 mm long) auricles at the base or the auricles absent. Inflorescences 4–10 cm long (excluding the awns), arched or nodding, disarticulating at the nodes of the axis, the joints shed as a unit with the attached spikelets. Spikelet clusters with the central spikelet fertile and the 2 lateral spikelets sterile and with reduced florets. Glumes 25–80 mm long (including the awns), narrow and awnlike throughout, slender, spreading or curved outward at maturity. Lemma of the fertile florets with the body 5.5–8.0 mm long, narrowly elliptic, tapered to an awn (10–)30–70 mm long, the awns slender, spreading or curved outward at maturity. Anthers 1.0–1.5 mm long. Fruits 3.0–3.5 mm long. 2n=14, 28, 42. May–October.

Scattered to common, mostly north of the Missouri River (northern U.S. south to North Carolina, Texas, and California; Canada; introduced in South America, Europe, and Asia). Banks of streams and rivers and edges of marshes; also pastures, crop fields, fallow fields, roadsides, railroads, and open, disturbed areas.

Steyermark (1963) noted, “The young inflorescence of this species sometimes takes on a beautiful rose‑purple color with a glistening effect in the sunlight.” It often occurs as dense populations along roadsides. The awns can injure the noses, eyes, mouths, and intestines of cattle and other species that attempt to graze on the plants, which makes the species undesirable as forage, except when very young.

For a discussion of a sterile hybrid between H. jubatum and Elymus trachycaulus (Link) Gould ex Shinners, see the treatment of that species.

The plant known as “bobtail barley” was reported as introduced in a St. Louis rail yard by Steyermark (1963, as H. jubatum var. caespitosum (Scribn.) Hitchc.). This taxon has a complicated nomenclatural and taxonomic history. Originally, it was described as a separate species, Hordeum caespitosum Scribn. Bowden (1962) reduced it to H. jubatum ssp. ¥intermedium Bowden and believed it to have arisen through hybridization between two other subspecies of H. jubatum (one of which was treated as a separate species, H. brachyantherum Nevski, by some other authors). Mitchell and Wilton (1964) summarized evidence for the hybrid nature of this taxon, which differs morphologically from H. jubatum in its shorter lemma awns (10–35 vs. 30–75 mm), but they considered it an interspecific hybrid between H. jubatum and H. brachyantherum (Hordeum ¥caespitosum). However, Baum (1980) completed a series of morphometric studies and concluded that plants in natural populations could be distinguished from artificial hybrids, and that these two groups should be treated as separate taxa (with the name H. caespitosum corresponding to wild plants). More recently, Baden and von Bothmer (1994) concurred that wild plants are a tetraploid (2n=28) taxon of hybrid origin, but they suggested that this is stabilized (self‑reproducing) and not separated sharply enough in morphology from typical H. jubatum to warrant separate species status. Thus, Baden and von Bothmer (1994) returned to Bowden’s (1962) treatment of the plants as a subspecies. The correct treatment of the single Missouri specimen is not at all clear, in spite of the considerable research cited above. For the present, it seems most convenient to treat it as a hybrid of uncertain origin, with H. jubatum as one of the parents.

 
 


 

 
 
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