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Published In: Bulletin de la Société Impériale des Naturalistes de Moscou 11: 356. 1838. (Bull. Soc. Imp. Naturalistes Moscou) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Synonym Text: P. mucronatum Royle ex Bab. in Trans. Linn. Soc. 18: 115. 1838; P. meyeri Steud., Nomen. Bot. Ed. 2, 2: 376. 1841; P. lemannianum Meisn. in DC., Prodr. 14: 91. 1856; P. himalaiense H. Gross. in Engler, Bot. Jahrb. 49: 343. 1913; P. mezianum H. Gross. l.c. 342.
Flower/Fruit: Fl. Per.: July-September.
Type: Type: Described from Talysch (LE).
Habitat: This Irano-Turanian element is more common in Eastern Irano-Turanian subregion; usually grows on dry rocky soil, dry ridges, hangs over cliffs. Very common in the dry inner zones, between 1200-4000 m.
Distribution: Distribution: Georgia (Caucasia), Turkmenistan, Pamir-Alai, Iran, Afghanistan, Pakistan and Kashmir (Western Himalayas).
Comment/Acknowledgements: A highly variable species particularly in size, branching and length of the internodes, shape of the leaves, apex and the length of ochreae. P. mezianum H. Gross is said to be characterized by having obovate-rounded leaves. However, even in the type specimen of P. mezianum H. Gross (Gilgit expedition, 1857, Giles s.n. (K!)) some of the leaves are not rounded but acute and even some of them are acuminate (spiny) therefore the former taxon seems to be a minor variant of P. paronychoides C. A. Meyer ex Hohen.
Map Location: A-6 Oihor Gol, Chitral, 36.4'N, 71o, 48', E, 11000 ft, 8.6.1958, S. A. Bowes Lyon 829 (BM); Barum Gol, Chitral, 4500 m, P. Wendelbo s.n. (BM); Lowari Pass, 9000 ft, Hariss 16588 (BM), Naltar, dry ridge, 10500 ft, R. R. Stewart 26263 (BM, RAW); Thui region c. 200 Km N. W. of Gilgit near the watershed separating the Gilgit Agency from Chitral, 4265 m, David J. Broadhead 79 (E); Bomboret, Chitral, 6800 ft, 35o.41'N, 71o.38'E, S. A. Bowes Lyon 642 (BM, E); Jamali, Gilgit, ± 11000 ft, Giles 457 (K); Arkari Gol between village Arkari and Avir, c. 3000 m A.Ghafoor & S. Omer 2802 (KUH); Chitral, Ziarat, 1908, S. M. Toppin 143 (K); Oihor Gol, 36°.4'N, 71°.48'E, fls. pink, alt. 11000 ft, S. A. Bowes Lyon 829 (E, K); Shogar Gol, Karimabad village area, W. of Tirichmir, 3000 m, A.Ghafoor & S. Omer 2928 p.p. (KUH); Barir village, 20 miles from Chitral on way to Bombrait, Sultanul Abedin 8143 (KUH); Girjar Kili, near Lowari Pass, M. Qaiser & A. Ghafoor 6549 (KUH); 8 miles from Lowari top towards Dir, 8000 ft, M. Qaiser & A. Ghafoor 1809 (KUH); B-7 Kalam, c. 2500 m. sandy-clay soil, A. Ghafoor & S. Omer 3366 (KUH); Kalam rocks 7000' Stewart 25093 (RAW); 3 miles from Naran on way to Saiful Muluk, Kamal Akhtar Malik & M. Qaiser 357 (KUH); Lake Saiful Muluk road, Hazara, ± 8000 ft, Anjum Amin, M. Arif & Sarfaraz Khan 63 (ISL); Naran, Mansehra, soil stony, M.N.Chaudhri et al. 1568 (ISL); Naran, Shaukat Ali 138 (RAW); Bahrein, Swat, 5000 ft., A. Rahman 162 (BM); Kaghan valley, 9000 ft, Inayat 20126 (K); Hazara dist., common on overgrazed flat ground, R. N. Parker 3409 (K); Kulalai, about Seven miles up Swat river from Bahrein, alt. 5500 ft. R. J. Rodin 5601 (K); Kaghan valley, Naran, 7800 ft, B. L. Burtt & M. Arshad Ali 825 (E); Plandran, Hazara, ± 4000 ft, Shaukat & Nasir 39 (ISL); Across river kunhar, fls. pink, M.N.Chaudhri et al. 1648 (ISL); ibid, June 1960, D. McVean s.n. (E); Naran, 2600 m, F. Schmid 275 (BM); Bahrein, 4500-5000 ft, Stewart & Rahman 25074 (BM, RAW). B-8 Gurikote, Astore, S. Omer & M. Qaiser 2452 (KUH); Gurikote, A. Kazmi s.n. (KUH); Shigar to Skardu, Schlagintweit 6179 (BM); Gurais, Kashmir, R.R. & I.D.Stewart 4392A (RAW); Rushkin Hill, Basho valley, Skardu, Katherin Hofmann et al. 29-1-3A (RAW); Above Shankargarh, Astor Dist., desert places, ± 10000 ft. R. R. Stewart 22271 (K); ibid, ± 9000 ft, March 1886, Giles s.n. (K); Above Rattu, upper Astor valley, 9500 ft, R.R. & I.D.Stewart 18804 (RAW); Bok, Zanskar, Kashmir, on dry slope, alt. 11,500 ft, fls. rose lavender, Walter Koelz 2947 (K); Skardu, Baltistan, 8000 ft, M. A. Siddiqi et al. 4123 (K, RAW); Zanskar, 2.6.1948, Herb. Hookerianum (K); Dah, Baltistan, 9-1000 m, in deserto, Aug. 1905, Fl., Meebold 3898 (K); Shagarthana valley, Baltistan, 10-11000 ft, 18.7.1892, J. F. Duthie s.n. (K); Suri, Kashmir alt. 10000 ft, B. B. Ostman 163 (K); Chatpani nala, 10-11000 ft, 27.8.1893, J. F. Duthie 13823 (E); Ladakh: Kharbu, common on dry hillside, 10000 ft, C. C. Burt 20(a) (K); Baltistan, Kashmir, 10-11000 ft, J. F. Duthie 13823 (BM); Zaskar, 12000 ft, Southampton University Botanical expedition 40 (BM); Dras, 10000 ft, 34.28N, 75,57E, J. D. A. Stainton, C. A. Mey 7829 (BM); B-9 Kargil Treaty road, Ladakh, 9000 ft, F. Ludlow & G. Sheriff 8346 (BM, E); Gilgit expedition, Giles s.n. (E); Deosai plains, Stewart 20012 (RAW); C-6 Kurram valley, Harsukh 14701 (K); Parachinar, Harsukh, 14729 (K); Parachinar, 33o56N', 70o04'E, 1500-1700 m, K. H. Rechinger 30892 (K); Kurram valley, Harsukh 15479 (K); Shalozan road, Parachinar, 5000 ft, Siddiqui & Nasir 6597 (RAW); D-5 Shenghar, Zhob district, Saood Omer & A. Ghafoor 1688 (KUH); near Karlachi, fls. pinkish, Y. Nasir & M. A. Siddiqi 8777 (KUH, RAW).


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Prostrate-subascending, perennial, often glabrous or sometimes scaberulous shrubby herb with thick woody rhizome. Stem woody, stunted or tufted, branched from base, branches spreading or tufted and prostrate, internodes very short, 2-4(-6) mm long, reddish brown, ± completely covered with leaves. Ochreae silvery hyaline, 4-6 mm long, ovate, lacerate, tubular, almost equalling or longer than the internodes. Leaves 7.5-15 x 0.7-1.5 mm, sessile, narrowly linear, acute or mucronate, margins recurved, scabrous. Flowers axillary, solitary, or 2 in each axil, sessile, enclosed in the ochreae. Ochreolae 1-2 mm long, ovate, lacerate, membranous. Tepals 5, pink, 1.5-2.0 x 0.75-1.0 mm, oblanceolate, acute-acuminate with slightly undulate margin. Stamens 4 (-6), filaments short, equal, dilated at the base. Ovary c. 1.0 mm long, lanceolate-ovate, trigonous with 3 styles and capitate stigma. Nuts trigonous, 1.5-2.0 x c. 0.7 mm broad. black, shining, glabrous.
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