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(Last Modified On 6/3/2013)
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(Last Modified On 6/3/2013)
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Species
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Vernonia canescens H.B.K.
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PlaceOfPublication
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Nov. Gen. Sp. P1. 4: 35, tab. 317. 1820.
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Synonym
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Verronia bullata Benth. in Orst., Vidensk. Middel. Dansk Naturhist. Foren. Kjobenhavn 1852: 67. 1852. Cacalia bullata (Benth.) Kuntze, Rev. Gen. P1. 969. 1891. Cacalia canescens (Benth.) Kuntze, Rev. Gen. P1. 969. 1891.
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Description
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Shrubs or coarse herbs to 3 m tall, erect, sparsely to many-branched; branches lignescent, striate, often densely pubescent above, glabrescent. Leaves short petiolate; petioles terete to canaliculate, 3-12 mm long; blades broadly lanceolate to ovate-elliptic, acute to acuminate at the apex, rounded to obtuse at the base, 4-12 cm long, 2-5 cm broad, marginally entire to remotely denticulate, often rugose to rarely bullate, scabrous to strigillose above, densely strigillose and glandular-punctate beneath, lateral veins ascending, 8-12 pairs. Inflorescences terminal in sparsely branched, short-pendunculate cymes, the cymes frequently arranged in large, spreading panicles or corymbs. Heads with 21-24 florets, sessile; involucre broadly campanulate, 4-6 mm high, 5-6-seriate, loosely imbricate, sparsely villous, the outer bracts triangular-subulate, spinose-tipped, the inner bracts lanceolate, acuminate at the apex, scarious; receptacle flat, alveolate, naked; corolla salverform, 3.5-5 mm long, 5-lobed, the lobes narrowly triangular, 2-2.5 mm long; stamens 5, anthers 2-2.2 mm long, apically acute, basally obtusely sagittate; styles 3-5.5 mm long, the 2 branches 1.5-2 mm long, antrorsely puberulous. Achenes narrowly turbinate, faintly ribbed, 1-2.5 mm long, antrorsely puberulent; pappus white, biseriate, the inner bristles linear, 3-5 mm long, the outer series subulate, 0.4-1.1 mm long.
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Habit
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Shrubs or coarse herbs
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Distribution
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ranges from southern Mexico southward throughout Central America to tropical South America. In Panama
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Note
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Vernonia canescens is a common small shrub. Specimens have been collected from Chiriqui Province with bullate leaves. Bentham segregated the bullate-leafed populations into V. bullata; however, the presence of a continuous gradation from the rugose to the bullate condition warrants the inclusion of V. bullata within the specific delimitations of V. canescens.
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Specimen
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CANAL ZONE: France Field, Blum & Dwyer 2117 (MO). Within 5 mi N of Gamboa gate, D'Arcy 6008 (MO). Albrook, Dwyer & Robyns 81 (MO); Dwyer 4469, 7318 (both MO). Chagres, Fendler 160 (MO). Ancon Hill, Greenman & Greenman 5124 (MO); Celestine 70 (NY, US); Williams 26 (NY). Near Miraflores, Greenman & Greenman 5202 (MO). Between Summit and Gamboa, Greenman & Greenman 5234 (MO). Las Cruces Trail, Hunter & Allen 690 (MO). Madden Forest Preserve, Lewis et al. 5364 (MO). Navy Reservation N of Gamboa, Robyns 65-44 (MO). Balboa, Standley 29314 (MO). 1 mi N of Summit, Tyson et al. 2769, 2774, 2779 (all MO). Curundui, Tyson 3477 (MO). BARRO COLORADO ISLAND: Busey 302 (MO); Croat 7079 (MO), 7219 (MO, NY), 7259, 7480, 8264 (all MO), 8392 (MO, NY), 9398 (MO); Shattuck 774 (MO); Wetmore & Abbe 208 (MO), Woodworth & Vestal 346 (MO). CHIRIQUI: Central Valley of Rio Chiriqui Viejo, Allen 1344 (MO, NY). NW of Boquete, Cerro Horqueta, Dwyer et al. 522 (F, GH, MO, NY, US). Rio Chiriqui Viejo Valley, near El Volcan, White 178 (MO). COCLE: Penonome, Williams 592 (NY). COLON: Santa Rita, G6mez-Pompa et al. 3215, 3294 (both MO). HERRERA: Vicinity of Oci', Stern et al. 1747 (MO). 10 mi S of Ocui, Tyson et al. 2802 (MO). LOS SANTOS: Loma Prieta, Duke 11835, 11890 (both MO). 16 mi S of Macaracas, at Quebrada Bejuco, Tyson et al. 2915 (MO). PANAMA: Taboga Island, Allen 129 (MO). Trail between Cafiazas and foot of Cordillera Central headwaters of Rio Cafiazas, Allen 187 (MO). Along road to Cerro Campana, Correa & Dressler 849 (MO). Cerro Azul, Duke 9342 (MO); Kant 45 (MO). Cerro Jefe, Duke 9385, 15240 (both MO). Isla del Rey, Duke 9523 (MO). Isla Saboga, Duke 10351 (MO). San Jose Island, Erlanson 71 (NY, US). 1 mi E of Charne, Lazor 2207 (MO). Cerro Campana, Porter et al. 4301, 4892 (both MO). Rio Piriquete, near Capira, Saldana 13 (MO). VERAGUAS: Carretera a -Santa Fe, Cisneros 10 (MO).
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