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Published In: Species Plantarum 2: 987. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 5/8/2022)
Acceptance : Accepted
Project Data     (Last Modified On 2/28/2022)
Contributor Text: Anjum Perveen & M. Qaiser
Contributor Institution: Centre for Plant Conservation, University of Karachi – PAKISTAN
Synonym Text:

Acanthoxanthium spinosum (L.) Fourr. in Ann. Soc. Linn. Lyon, Ser. 2, 17: 110. 1869; X. spinosum L. var. inerme Bel, Rev. Bot. Bull. Mens. 11: 481. 1891.

Flower/Fruit:

Fl. Per.: July – August                                          Engl. Vern.: Bathurst burr

Type:

Lectotype: Habitat in Lusitania, Herb. Linn. No. 1113.3 (LINN) (designated by Wijnands, Bot. Commelins: 87. 1983).

Distribution:

Native of North and South America, widely naturalized in Old and New World.

Comment/Acknowledgements:

This South American species has spread widely in many regions of the world, probably via its spiked burs which attach to animals and clothing or as a contaminant of hay or other products. It is now established as a common and one of the most noxious agricultural and pasture weeds. It produces huge amount of cypselas which germinate easily. X. spinosum can quickly dominate large areas, outnumbering crops, forage plants and native flora.

Map Location:

D–5 Zhob Appozai, 24.10.2019, Nazar Khan Mandokhel 05 (KUH); Zhob city, 24.10.2019, Kamran Ishaq Bahadikhel 06 (KUH).


 

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Annual, 25–100 cm tall, branched herbs, with rigid, nodal trifid spines at the base of leaves, spines yellow, 1.5–2.5 cm long. Stem pubescent, younger part of stem green, woolly. Leaves simple, whorled or opposite, linear to lanceolate, upper leaves undivided, lower usually 3–lobed, on 3–10 mm long petiole, lamina 2.5–3.5 cm long, discolourous, adaxial surface dark green, ± pubescent, white tomentose abaxially. Male capitula small, 5–6 mm across, discoid, green, 5–7 flowered. Phyllaries 2–seriate, outer ones 6–7, lanceolate, 1–1.5 mm long, green, pubescent, acute, inner ones 6–7, lanceolate to ovate, 1.5–2 mm long, green, pubescent acute. Receptacle paleate, paleae oblanceolate to spathulate, 2–2.5 mm long, hairy. Functionally staminate florets 2.8–3 mm long with rudimentary stigma, style and ovary. Corolla tubular, creamy yellow, 5–lobed, anthers exerted. Female capitula usually solitary, 6–15 mm long, 2 – flowered, phyllaries 2–seriate, outer phyllaries 5–7, green,  4–6 mm long, inner ones united, enclosing the female florets, spiny, spines numerous, 1.5–2 mm long, reaching upto ca 2.5 mm long on maturity, golden brown, 2–beaked. Female florets 4–6 mm with bifid, exserted stigma.  Cypselas enclosed by hooked, spiny phyllaries (burs), burs 12–15 mm long, ca 5 mm wide, cypselas black, ± glabrous, epappose, sometimes with occasional bristles.

 
 
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