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(Last Modified On 6/14/2013)
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(Last Modified On 6/14/2013)
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Species
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Clibadium pittieri Greenman
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PlaceOfPublication
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Proc. Amer. Acad. Arts 39: 98. 1903.
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Note
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TYPE: Costa Rica, Limon, La Florida, 80 m, Jul 1897, Pittier 11290 (CR, GH, syntypes, not seen).
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Synonym
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Clibadium pittieri f. phrixium Greenman, Proc. Amer. Acad. Arts 40: 38. 1904. SYNTYPES: Costa Rica, "Buissons 'a Tuis," 650 m, Dec 1897, Tonduz 11479 (CR, GH, not seen, US). "Confluent du Puerto Viejo et du Sarapiqui," Jan 1893, Biolley 7399 (CR, GH, not seen).
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Description
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Shrub 1-3.5 m tall; stems antrorsely strigose to hispidulous with hairs 0.1-1 mm long. Leaves narrowly ovate to ovate; blades 8-19.5 cm long, 4-10 cm wide, acuminate, the base attenuate, the margin serrate, both surfaces weakly or moderately strigose to hispidulous with hairs to 0.7 mm long; petioles 1.8-7 cm long, to 1 mm diam., strigose to hispidulous with hairs to 0.5 mm long. Inrflores- cence a corymbose panicle with 8-26 heads; peduncles 1-2 mm long. Heads radiate; involucre cupulate, 5-6 mm across, the bracts 6-8, narrowly ovate to ovate, 2.6-3.2 mm long, 1.3-2.1 mm wide, abaxially strigose toward the apex with hairs 0.2 mm long, ciliate; paleas subtending both ray and disc florets, lanceolate, scarious, 2-3 mm long, 0.3-0.7 mm wide, on abaxial surface strigillose toward apex with hairs 0.3 mm long; ray florets 22-26, the corollas 1 mm long, 0.3 mm diam., the 4 irregular lobes 0.3 mm long, the styles 1.7 mm long with branches 0.8 mm long; disc florets 5-10, the corollas 1.6 mm long, the throat 1 mm long, 0.6 mm diam., the lobes 0.6 mm long, weakly pubescent near apex, the tube 0.3 mm diam., the anthers 1 mm long, the style 2.7 mm long, the ovary sterile, 1.7 mm long, 0.2 mm diam., pilose (especially near apex) with hairs to 0.6 mm long. Achenes 2 mm long, 1.8 mm diam., pubescent on upper 1/? with hairs 0.1 mm long; pappus absent. Chromosome number unknown.
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Habit
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Shrub
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Note
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Within Clibadium pittieri, two different types of pubescence are found on the young stems and peduncles. In some plants, the vesture is strigose (Duke & Elias 13778), and in others it is hispidulous (Wilbur & Teeri 13423). Despite this obvious difference, the populations of each type are not recognized as varieties here because no geographic separation prevails and because one collection (Lewis et al. 975) is clearly intermediate in these pubescence features. One could recognize these differences by describing and naming formas (as Greenman did in recognizing forma phrixium for the plants with hispidulous vesture), but I do not subscribe to this policy. In Panama this species flowers in all months except June to August.
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Specimen
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BOCAS DEL TORO: Changuinola Valley, Dunlap 283 (F, US). 10-15 mi S from mouth of Changuinola River, Lewis et al. 975 (GH, MO, UC, US). Old Bank Island, near Chiriqui Lagoon, von Wedel 2161 (MO). CANAL ZONE: Ca. 8.5 mi NW of Gamboa, Wilbur & Teeri 13423 (DUKE). COLON: Near Guasimo on Rio Miguel de la Borda, Croat 10005 (MO). Santa Rita Ridge, Porter et al. 4785 (MO). DARIEN: Camp Summit, Blackwell et al. 3667 (MO). Cerro Pirre, Bristan 538 (MO). Near Boca Quebrada Venado, Rio Tuqueza, Bristan 1110 (OS). Cerro Pirre, Duke 6088 (MO). Camp Summit, ca. 1200 ft, Duke 15487 (OS). Cerro Pirre, 2500-4500 ft, Duke & Elias 13778 (MO). Camp Summit, adjacent to Darien-San Blas border, 1000-1200, ft. Oliver et al. 3667 (MO). PANAMA': 5 km NE of Altos de Pacora, Busey 825 (MO). 3; mi above Interam. Hwy. on road to Cerro CAmpana, Croat 12070 (MO). Cerro Campana along trail to Summit, Croat 17175 (MO). Headwaters of Rio Corso (off Rio Pacora), ca. 50() m, Duke 11918 (MO, OS). Cerro jefe, 3100 ft. Dwyer et al. 9496 (MO, NY). 5 mi SW of Cerro Brewster, ca. 1000 ft,,Lewis et al. 3434 (MO'). E slope Cerro Jefe, 2700 ft. Tyson 3422 (MO ). VERAGUAS: 5; mi NW of Santa Fe, 700-1200 m, Croat 23190 (MO).
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