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(Last Modified On 9/18/2013)
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(Last Modified On 9/18/2013)
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Species
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Tocoyena pittieri (Standley) Standley
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PlaceOfPublication
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Contr. Arnold Arbor. 5: 151. 1933.
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Synonym
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Posoqueria pittieri Standley, J. Wash. Acad. Sci. 18: 167. 1928. TYPE: Panama, Pittier 6635 (US, holotype). Tocoyena cuatrecasasii Steyermark, Act. Biol. Venez. 4(1): 113. 1964. TYPE: Colombia, Cuatrecasas 1918.
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Description
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Trees to 10 m tall or much taller (fide Croat), the branchlets terete, smooth but often nodose, glabrous, the nodes usually 2-3 cm apart. Leaves elliptic, to 32 cm long, to 17 cm wide, widely deltoid to somewhat rounded at the apex, shortly acuminate, the acumen to 1 cm long, ultimately obtuse, cuneate toward the base, often somewhat inequilateral, the lateral veins ca. 10, arcuate, the lesser veins slender, irregular and spreading, pinnatiform, plane, stiffly chartaceous, black brown when dry, glabrous except the minutely pubescent axils of the prin- cipal veins beneath, minutely papillate under magnification, lightly marcescent above; petioles to 3 cm long, ca. 0.25 cm wide, glabrous; stipules free, triangular, 5-10 mm long. Inflorescences cymose paniculate, resembling a candelabra, to 13 cm long, ca. 11.5 cm wide, the flowers numerous, erect; bracts and bracteoles triangularly subulate, 0.5-1 mm long. Flowers pedicellate, the pedicels 1-3 mm long; hypanthium oblong, to 4 mm long, truncate, glabrous, the calycine cup ca. 1 mm long, eglandular within, the teeth 5, widely triangularly subulate, to 0.5 mm long, glabrous; corolla yellow, the tube narrowly cylindrical, 8-12 cm long, to 0.6 cm wide, thickly carnose, glabrous outside, within white villous at the base of the tube, the lobes 5, forming an ovate rotund mass in bud, golden farinose outside, ovate elliptic, obovate or elliptic rotund, 1-1.5 cm long, glabrous; sta- mens 5, exserted, sessile, the anthers elliptic, ca. 6 mm long, ca. 2 mm wide,
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Habit
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Trees
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Description
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attached at the corolla mouth; style slender, ca. 0.6 mm wide, the stigmas ovate- lanceolate, crassate, ca. 5 mm long. Fruits sessile, solitary, subrotund, 6-10 cm in diam., woody when dry, the wall thick, to 1.3 cm wide, drying black outside, tan within, the pulp blue black when dry, smooth externally except longitudinally costate, the ribs 10-15, well spaced, slender or thick, prominent but not ridge- like, often evanescing along their length, the surface often marked by lenticels; seeds flat, ovate trapeziform, to 2 cm long, to 1.3 cm wide, slimy to touch.
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Distribution
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known only from Panama.
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Note
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Kirkbride and Duke (1368) note: "anthers white, reflexed with the petals; seeds embedded in a brownish- black pulp with somewhat the odor and taste of molasses."
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Specimen
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CANAL ZONE: Summit Garden, Lindsay 322 (MO, US). Trinidad River near Hydrographic station, Pittier 6635 (US). BARRO COLORADO ISLNAD: Aviles 14 (MO); Croat 4636, 8529 (both MO); Hladik 138 (MO), 299 (MO, US); Shattuck 671 (MO); Starry 131 (MO); Wetmore & Abbe 197 (MO); Wood- worth & Vestal Zetek 3536 (MO). DARIEN: Rio Ucurganti, Bristan 1185 (MO). La Boca de Pirre, Bristan 1246 (MO). Rio Yape, Bristan 1399 (MO). Trail from Cerro Mali to Rio Pucro, 650-1200 m, Gentry et al. 16970 (MO). Rio Balsa near Rio Cuasi, Kirkbride & Duke 1386 (MO). Rio Balsa near Cerro Campamento, S. Cerro Pirre cloud forest, Duke 15999 (MO). PANAMA: La Cuesta, Panama City, Holdridge 5614 (MO).
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