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Faramea trinervia K. Schum. & Donn. Sm. Search in The Plant ListSearch in IPNISearch in Australian Plant Name IndexSearch in NYBG Virtual HerbariumSearch in Muséum national d'Histoire naturelleSearch in Type Specimen Register of the U.S. National HerbariumSearch in Virtual Herbaria AustriaSearch in JSTOR Plant ScienceSearch in SEINetSearch in African Plants Database at Geneva Botanical GardenAfrican Plants, Senckenberg Photo GallerySearch in Flora do Brasil 2020Search in Reflora - Virtual HerbariumSearch in Living Collections Decrease font Increase font Restore font

Published In: Botanical Gazette 31(2): 115. 1901. (23 Feb 1901) (Bot. Gaz.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 11/27/2017)
Acceptance : Accepted
Project Data     (Last Modified On 11/28/2017)

This species is characterized by its medium-sized, elliptic to oblanceolate subsessile to shortly petiolate leaves, shortly sheathing and shortly aristate stipules, terminal, paniculiform, blue inflorescences, short dentate calyx limbs, somewhat small blue corollas with the tubes longer than the lobes, and medium-sized fruits that are oblate and laterally flattened. The leaf venation has the characteristic arrangement found in many Faramea species, with the secondary veins straight and connecting to form an equally well developed, generally straight submarginal vein and one or more well developed intersecondary veins present between the pairs of secondary veins. The leaves of Faramea trinervia are weakly to markedly bullate in life and usually have at least the secondary and submarginal veins impressed, but often are flat on dried specimens. The inflorescences are pedunculate and branched to several orders, with pedicellate flowers.

Faramea trinervia has been distinguished from Faramea suerrensis by subsessile leaves with cordate bases that clasp the stems, vs. leaves with acute to rounded bases and well developed petioles. The plants that agree with this diagnosis of Faramea trinervia are found in a somewhat small area, and show a notable range of variation in leaf size, number of flowers in the inflorescence, diameter of the corolla tubes (at least on dried specimens), and petiole development; this variation is similar to that found in Faramea suerrensis. Leaf blade size varies widely in Faramea trinervia, by more than 300%, from 8-10 x 2-2.5 (e.g., Robles & Chacón 2692) to 37 x 15 cm (e.g., Batista et al. 1483). Development of the petiolate and the shape of the leaf base also vary notably, from reduced to up to 1 cm long and truncate to deeply cordate, respectively, and their variation does not seem to be closely correlated. This means some plants are intermediate between Faramea trinervia and Faramea suerrensis. These variable, often intermediate plants that agree with Faramea trinervia could represent variation within a population that is not completely distinct, due to either ongoing differentation (in an area with some locally endemic Rubiaceae species) or hybridization of species that are newly sympatric; field study will be needed to evaluate this situation. These two species are separated here until further study clarifies their status, and if they are later combined, then Faramea trinervia is the older and thus correct name for the combined species.

Plants similar to Faramea trinervia and Faramea suerrensis with shortly developed petioles, relatively quite narrow leaves with cordulate to truncate bases were separated as Faramea scalaris by Lorence et al. (2012). Continuous variation now is documented that links these with plants that match the type Faramea trinervia, and these are combined here.

Distribution: Wet forest at 10-1200 m in east-central Costa Rica through western Panama.



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