Accepted By:
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AFPD. 2008. African Flowering Plants Database - Base de Donnees des Plantes a Fleurs D'Afrique.
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Bridson, D.M. & B. Verdcourt. 1988. Rubiaceae (part 2). 415–747. In W. B. Turrill & R. M. Polhill (ed.) Fl. Trop. E. Africa. A. A. Balkema, Rotterdam.
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Hiern, W. P. 1878. On the peculiarities and distribution of Rubiaceae in tropical Africa. J. Linn. Soc., Bot. 16: 248–280.
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Schatz, G. E., S. Andriambololonera, P. P. Lowry II, P. B. Phillipson, M. Rabarimanarivo, J. I. Raharilala, F. A. Rajaonary, N. Rakotonirina, R. H. Ramananjanahary, B. Ramandimbisoa, A. Randrianasolo, N. Ravololomanana, C. M. Taylor & J. C. Brinda. 2023. Catalogue of the Plants of Madagascar.
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