Accepted By:
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AFPD. 2008. African Flowering Plants Database - Base de Donnees des Plantes a Fleurs D'Afrique.
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Kårehed, J. & B. Bremer. 2007. The systematics of Knoxieae (Rubiaceae)-molecular data and their taxonomic consequences. Taxon 56(4): 1051-1076.
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Schatz, G. E., S. Andriambololonera, P. P. Lowry II, P. B. Phillipson, M. Rabarimanarivo, J. I. Raharilala, F. A. Rajaonary, N. Rakotonirina, R. H. Ramananjanahary, B. Ramandimbisoa, A. Randrianasolo, N. Ravololomanana, C. M. Taylor & J. C. Brinda. 2024. Catalogue of the Plants of Madagascar.
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Taylor, C. M. 2010-2025. Studies in Tropical Rubiaceae.
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Verdcourt, B. 1975. Studies in the Rubiaceae-Rubioideae for the 'Flora of Tropical East Africa': I. Kew Bull. 30(2): 247–323.
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