Accepted By:
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Lorence, D. H. 1999. A nomenclator of Mexican and Central American Rubiaceae. Monogr. Syst. Bot. Missouri Bot. Gard. 73: 1–177.
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Standley, P. C. & L. O. Williams. 1975. Rubiaceae. En: Standley, P.C. & Williams, L.O. (eds.), Flora of Guatemala - Part XI. Fieldiana, Bot. 24(11): 1–274.
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