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(Last Modified On 10/31/2013)
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Species
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Rondeletia ginetteae Lorence
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Note
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TYPE: Mexico. Oaxaca: Distrito de Tuxtepec, Ruta 175 Tuxtepec to Oaxaca, ca. 10 km S of Valle Nacional, Sierra de Juarez, ca. 750 m, 6 Dec. 1980, D. H. Lorence & R. Cedillo T. 2973 (holotype, MEXU, photo PTBG; isotype, F, photo PTBG).
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Description
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Shrubs 1-2 m tall, the young twigs densely hir- sutulous to hirsute, the trichomes brown or purple, to 1 mm long, the twigs dark brown, 1-2 mm diam., the internodes 1-10 cm long. Leaves opposite, those of a pair at a node subequal or unequal, one oc- casionally up to 6 times larger than the other, shortly petiolate or subsessile; petioles 1.5-3 mm long, 1- 1.5 mm diam., densely hirsutulous-hirsute; lamina elliptic or obovate-elliptic, (5-)9-23 x (2-)4-11 cm, often somewhat falcate, chartaceous, drying green or pale brownish green, slightly discolorous, adaxially hirsutulous, especially along the costa and 2? veins, abaxially more densely whitish hirsutulous or villosulous, the trichomes longer and denser along the costa and veins, the base obtuse, rounded, or subcordate, the apex shortly acuminate, the acumen 1-3 cm long, often falcate, the 2? veins 7-10 pairs, camptodromous or weakly brochidodromous, the 2?(- 3?) veins depressed adaxially, the venation very prominent and visible to 4? on both surfaces, finely reticulate, the margin ciliolate; stipules erect, ellip- tic-lanceolate, acuminate, 8-13 x 1.5-3 mm, thin, brown, externally strigose especially basally and along margins, internally densely short strigose, with a ring of white trichomes 1-2 mm long basally. In- florescence terminal, cymose, thyrsiform-spiciform, 5.5-8 x 3-4 cm including the corollas, the pe- duncle 1-4.5 cm long, the flowers numerous, the axes hirtellous-tomentose, brownish purple, the pri- mary branches 3-6 pairs, crowded along the rachis, bearing sessile or subsessile glomerulate cymules of 4-6 flowers, subtended by thin, 2-3-lobed bracte- oles 4-5 mm long; flowers 4-merous, sessile, the hypanthium obconic, 1-1.5 x 1 mm, the thin calyx purplish brown, externally densely arachnoid to- 141 mentose, internally strigillose, the calyx cup 1-1.5 mm long, the lobes subequal, ovate or elliptic, 4-7 x 1.5-2.5 mm, venose, apically acuminate, inter- nally with 2 or 3 reddish colleters in each sinus; corolla reddish purple in bud, at anthesis salverform, tube externally densely arachnoid tomentose, brown- ish pink, 13-15 mm long, 2-2.5 mm diam., inter- nally villosulous basally, the lobes ovate or rounded, obtuse, 2.5-3.5 mm long and wide, spreading, ex- ternally reddish purple basally, tomentose, internally glabrous, white, the margin crisped; stamens 4, in short-styled flowers sessile, attached 1-1.5 mm be- low throat, anthers 2-2.5 mm long, ellipsoid, tips exserted; style in short-styled flowers glabrous, in- cluded, 8 mm long, the ellipsoid stigmas 2 mm long. Capsule subglobose-ellipsoid, 4-5 x 4-5 mm, hir- tellous, crowned by the persistent calyx lobes, de- hiscence loculicidal then septicidal; seeds angulate, 0.4-0.7 x 0.3-0.4 mm, the testa reticulate, dark brown.
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Habit
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Shrubs
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Note
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Known only from the lower north- ern slopes of the Sierra de Juarez in Oaxaca, Mexico. Rondeletia ginetteae occurs in lower montane tropical evergreen rainforest with Termin- alia amazonia, Dussia mexicana, and Swartzia from about 150 to 900 m (not 1,600 m as erro- neously indicated on the isotype). The type was collected from a wet, vertical rock face in a shady arroyo. Flowers were collected in December and January, and fruits were found in April.
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Specimen
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specimens examined. MEXICO. OAXACA: District of Tuxtepec, 5 km N of Valle Nacional, 150 m, Breedlove & Almeda 56774 (CAS); Distrito de Ixtlan, Mun. de Comaltepec, Sierra de Juarez, Ruta 175 a 2.5 km al NE de Puerto Eligio, 900 m, Cedillo T. & Lorence 2395 (BM, MEXU, MO, PTBG).
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Note
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Because of its somewhat elongate, spiciform-thyr- siform inflorescence and tetramerous flowers, Ron- deletia ginetteae keys out to the Laniforae group in Standley (1918). It is, however, clearly most closely related to R. acuminata (Oerst. ex Standl.) Lorence & Castillo-Campos and also to R. macro- calyx Standl. & Steyerm., both of which obviously belong in Standley's Calycosae group. This again underscores the artificiality of these groups. Al- though it occurs in the same region as the latter two species, R. ginetteae is readily distinguished by its elongate, spiciform inflorescence with flowers in paired, densely glomerulate cymules, smaller sub- equal calyx lobes, arachnoid-tomentose corolla with a brownish pink tube and externally reddish purple lobes toward the base, and smaller, subglobose cap- sules. It is with pleasure that I name this species for my wife, Ginette, frequent field companion and dearest friend.
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