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Project Name Data (Last Modified On 1/30/2013)

Flora Data (Last Modified On 1/30/2013)
PlaceOfPublication Sp. P1. 444. 1753
Reference Lay, Ann. Missouri Bot. Gard. 37: 378, fig. 12. 1950.
Synonym Triumfetta heterophylla Lam., Encycl. Meth. Bot. 3: 420. 1789. Triumfetta sinuosa Miq., Linnaea 22: 466. 1849. Triumfetta Hostmanni Miq., loc. cit. 466. 1849. Triumfetta Berterii Turcz., Bull. Soc. Imp. Nat. Moscou 31 (1): 227. 1858. Triumfetta Salzmanni Turcz., loc. cit. 31 (1): 229. 1858. Triumfetta quinqueloba Turcz., loc. cit. 36(1): 574. 1863.
Description Shrub ca. 0.60-2.50 m. high, the branches and inflorescence-axes ferruginous- pubescent. Leaves with a petiole ca. 2-8 cm. long or sometimes slightly more, ferruginous-arachnoid, the blade broadly ovate, sometimes pandurate, often 3- to 5-lobate, the lobe tips, at least the terminal, acuminate, ca. 8-10 cm. long and 7-9 cm. wide, the margins irregularly serrate, the serrations usually glandular, the up- per surface pubescent, the lower one arachnoid to pubescent, 3- to 5-palminerved, the nervation slightly prominent beneath. Inflorescences axillary, the cymes of 2, rarely 3 cymules, generally condensed, the bracts ovate to more or less elliptic, rarely obscurely 3-lobate, the peduncles and pedicels short, 1-2 mm. long, sparsely hirsute. Flowers 4.5-5.5 mm. long, hermaphrodite, the sepals 5, narrowly oblong, acute, with a very short apical appendage ca. 0.5 mm. long, ca. 4.5-5.5 mm. long and 0.5 mm. wide, fleshy, sparsely hirsute without, glabrous within; petals absent; gonophore and urceolus obsolete, the glands absent; stamens (5-) 10(-15), the filaments ca. 3-4 mm. long, glabrous, the anthers ca. 0.5 mm. long; ovary subor- bicular, ca. 0.6-0.8 mm. in diam., covered with numerous uncinate spinules; style ca. 3-3.5 mm. long, the stigma obscurely 2- to 3-parted. Fruit broadly elliptic, the body 3-4 mm. long and 2.5-3 mm. broad, lightly stellate-pubescent, the spines about 50, ca. 1.8-2 mm. long, retrorsely pilosulose, the spinules uncinate, 2- or 3- celled, each cell 1-seeded; seeds pyriform, ca. 1.5-2 mm. long and 1-1.5 mm. wide.
Habit Shrub
Distribution Common throughout tropical America; usually in dry open habitats, often in secondary formations
Common cadillo cepa de caballo
Specimen BOCAS DEL TORO: Changuinola Valley, Dunlap 122, 372, 433; vicinity of Chiriqui La- goon, Old Bank Island, von Wedel 1883, 1949, 2005; Water Valley, von Wedel 1710, 1779, 1825, 1847, 1854; Darkland, von Wedel 2618. CANAL ZONE: Chagres, Fendler 172; vicinity of Fort Sherman, Standley 30993; near Fort Randolph, Standley 28638; Mount Hope Cemetery, Standley 28828; Gatun, Bro. Heriberto 124, Hayes 370, 991; hills west of the Canal, near Gatun, Standley 27275; around the Agua Clara Reservoir, near Gatun, alt. 20-30 in., Maxon 4660; Barro Colorado Island, Shattuck 657, 728, Wetmore & Abbe 88, 174, Wilson 41, Wood- worth & Vestal 627; Frijoles, Standley 27638; Darien Station, Standley 31523; Gamboa, Stand- ley 28533; Obispo, Standley 31735; Summit, Standley 26941; around Culebra, alt. 50-150 m., Pittier 2145; vicinity of Miraflores Lake, P.White 279; Corozal, Standley 27392; vicinity of Corozal, Piper 5302, 5308; Ancon Hill, 600 ft., R. S. Williams 21; Balboa, Standley 27124, 32110. COCLk: Aguadulce, near sea level, Pittier 4961; El Valle de Ant6n, trails near Finca Tomas Arias, 600 m., Allen 4229. COLON: between France Field, Canal Zone and Catival, Standley 30242, 30423; Rio Viejo, vicinity of Puerto Pilon, 10 m., Allen 4100; Juan Mina plantation, Rio Chagres, Region above Gamboa, 25 m., Allen 4125. DARIEN: trail between CanA and Boca de Cupe, vicinity of El Real, along road to Rio Pirre, Stern, Chambers, Dwyer, Ebinger 610; locality unknown, Macbride 2686. HERRERA: vicinity of Ocd, 100 m., Allen 4056. PANAMA: vicinity of San Carlos, alt. 0-10 m., Allen 1138; between Capira and Potrero, 80-130 in., Dodge & Hunter 8626; along the Corozal Road, near Panama, Standley 26792; Sabanas northeast of Panama City, Bro. Paul 184; Paitilla, Bro. Heriberto 236; Punta Paitilla, Standley 30805; Bella Vista, Standley 25339; Agricultural Experiment Station at Matias Hernandez, Pittier 6926; Rio Tapia, Standley 26126, 28057; near the big swamp east of the Rio Tocumen, Standley 26698; Tocumen area, Dwyer 4884; Brujas, Mell s.n.; Taboga Island, Standley 27043, 27989. VERAGUAS: vicinity of Santiago, alt. ca. 50 m., Allen 1084. PROVINCE UNKNOWN: Carleton 122, Hayes 321.
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