Plants low, loosely to densely tufted. Stems generally with a thin-walled epidermis and usually a central strand. Propagula often present in leaf axils. Leaves elliptic to spatulate; margins plane to broadly incurved (or sometimes narrowly recurved below), entire or dentate; costa ending below the apex, percurrent, or ending in an apiculus, with 2 stereid bands; upper cells usually mammillose-bulging on the upper surface and slightly convex on the lower, occasionally also papillose; basal cells differentiated. Perichaetial leaves shorter than stem leaves. Setae terminal, elongate; capsules erect, cylindric; operculum conic; peristome none. Calyptra smooth, naked.