Plants medium-sized to large, in usually flattened, pale green to yellowish green masses. Primary stems creeping; rhizoids smooth, reddish brown, not or irregularly branched, densely present on stolons and secondary stems and branches when in contact with substrate; leaves erect, small. Sec-ondary stems erect, horizontal or pendent, not stipitate, irregularly branched, complanate-foliate; sec-ondary stems in cross section with sclerodermis and enlarged, thick- to firm-walled cortical cells, cen-tral strand absent; paraphyllia present or absent; pseudoparaphyllia foliose. Secondary stem and branch leaves erect-spreading or spreading, undulate (plane in species outside the region), oblong-ovate, oblong-lanceolate, or ovate-lanceolate, symmetric or asymmetric, acute to broadly acute, long-or short-decurrent, subauriculate, rounded or straight to the insertion; margins subentire, serrulate or denticulate, broadly inflexed on one side below, plane or narrowly recurved on other side; costae faint, short and double with one fork usually longer than the other (long and single outside the region); cells smooth, apical cells hexagonal, rhomboidal or fusiform, firm- to thick-walled; basal cells linear-fusiform, thick-walled, often porose; alar cells hyaline or reddish yellow, not or moderately devel-oped, cells across the insertions yellow. Autoicous. Setae smooth, yellow to yellow red. Capsules immersed or exserted, erect, cylindric, symmetric; exothecial cells not strongly differentiated at mouth, those below short-rectangular, firm- or thick-walled; stomata present or absent; opercula conic-rostrate; peristome diplolepideous; exostome teeth whitish yellow, dorsal (outer) surface cross-striate at base, trabeculae and median lines faintly developed, ventral (inner) surface smooth to weakly papillose, trabeculae well developed; endostome light yellow, lightly papillose, basal mem-branes variously developed, segments perforate, not keeled, more strongly thickened on the ventral (inner) surface than on the dorsal (outer) surface, cilia rudimentary or absent. Spores papillose. Ca-lyptrae cucullate, naked, smooth.