Plants robust, in dull, green or yellow tufts. Stems irregularly branched, not or sparsely radiculose. Leaves curled and contorted when dry, squarrose-recurved from an erect, sheathing base when moist, oblong-lanceolate to long-lanceolate from an oblong or ovate base, broadly acuminate and sharply acute; margins undulate, denticulate above the middle; costa strong, percurrent or short-excurrent, prominently rounded at back, with 4-6 median guide cells and 2 well-developed stereid bands, covered ventrally by short, papillose cells, dorsally by elongate, smooth cells; upper cells convex, subquadrate to short-rectangular, pluripapillose; basal cells large, rectangular to rhomboid, thick-walled within, lax, thin-walled, hyaline and smooth at the margins and extending well up the leaf as a distinct border. Dioicous. Inflorescences lateral. Perichaetial leaves sheathing, shorter and narrower than stem leaves. Setae elongate; capsules cylindric; annulus persistent; operculum high-conic or conic-rostrate; peristome divided to the base into 32 orange-red, branched-spiculose filaments twisted counterclockwise in 1 turn. Calyptrae cucullate, smooth, naked.