Plants small to large, dull or glossy, golden to yellowish green. Stems prostrate, simple or irreg-ularly pinnately, or tripinnately branched, at times filiform-attenuate, branches sometimes easily detached; stems in cross section with epidermis of small, thick-walled cells, central strand absent; pa-raphyllia absent; pseudoparaphyllia absent, scale leaves present; rhizoids from small clusters of ini-tials abaxial to the leaf insertions, smooth, not or sparsely and irregularly branched. Stem and branch leaves similar or dimorphic, oval, ovate, oblong, oblong-ovate, oblong-lanceolate, or lingulate, plane to concave, at times undulate, slightly asymmetric, obtuse, acute, or acuminate, not or shortly decur-rent; margins serrulate to coarsely serrate, plane or inflexed, often incurved on one side at base; costae short and double, extending to 1/4–1/2 the leaf length; cells smooth or prorate on dorsal surface, firm-walled, median cells linear, extreme apical cells shorter; alar cells more or less differentiated, of sev-eral quadrate to short-rectangular cells. Asexual gemmae sometimes present on stems. Dioicous. Setae elongate, smooth below, roughened to papillose above. Capsules erect to suberect, cylindric to oblong-ovate, symmetric, mammillose at the neck, densely spinose above; exothecial cells thick-walled, rectangular, not collenchymatous; stomata present; opercula conic, apiculate to long-rostrate; annuli well developed; peristome diplolepideous; exostome teeth narrowly triangular, dorsal (outer) surface smooth, faintly papillose, or weakly cross-striate at base, median lines faint, ventral (inner) surface smooth or papillose; endostome basal membranes low, smooth or faintly papillose, segments narrow, keeled, perforate, lightly papillose, cilia absent. Calyptrae cucullate, naked, smooth.