Syntrichia Brid., J. Bot. (Schrader) 1(2): 299.
Plants small, medium- to large-sized, scattered and gregarious or forming tufts or mats, yellowish green to reddish green, corticolous, saxicolous or terricolous. Stems smooth, erect, irregularly branched, red, central strand present, sclerodermis present or absent, hyalodermis present or absent; rhizoids scattered or densely clustered at base of stem, smooth, densely branched. Leaves evenly spaced, fragile or firm, oblong-lingulate, lingulate, spathulate, ovate-lanceolate, erect or clasping at base, appressed-flexuose, twisted and crisped or spirally twisted around stem when dry, erect-spreading to squarrose-recurved when wet; apices acute, obtuse or cucullate, apiculate, mucronate, or tapered to a long, reddish, sparsely spinulose awn; laminae unistratose or bistratose; margins entire or crenulate, at times bordered by thick-walled, colored cells, plane above, recurved below; costa subpercurrent, percurrent, excurrent, to long-excurrent, guide cells and single (dorsal) stereid or substereid band present, ventral surface layer of enlarged cells present; upper cells irregularly rounded-quadrate, hexagonal, short-rectangular, oblate, or triangular, firm-walled, irregularly thickened to collenchymatous, pluripapillose, by low, o-shaped, or high, branched, c-shaped papillae, basal cells enlarged, rectangular, smooth, alar cells not differentiated. At times with asexual gemmae. Dioicous. Setae elongate, smooth. Capsules exserted, cylindrical, erect; stomata at base; annuli of 2–3 rows of vesiculose cells; opercula rostrate; peristome irregularly spirally-twisted, basal membrane short. Spores granulate. Calyptra cucullate, smooth.