Plants up to 2.5 cm high, light- to dark-green, often ± glaucous, sometimes red-tomentose below. Stems seldom branched, with a central strand. Leaves of nearly equal size throughout, when dry spreading from the insertion and incurved, slightly twisted to crisped or catenulate, and subtubulose above, when moist spreading, oblong- to linear-lanceolate, the base scarcely differentiated to shortly ovate, not sheathing; margins plane to incurved, entire to dentate; costa ending below the apex to stoutly excurrent, bulging at back, covered ventrally by quadrate to elongate cells, the dorsal epidermis sometimes differentiated, with 2 usually strong stereid bands and 1 row of guide cells; upper cells bulging, subquadrate, hexagonal, or elliptic, thick-walled with papillae usually massive, bifid to multifid, usually centered over the lumen, with 4-6 salients per cell; basal cells differentiated in a small yellowish to hyaline area, rectangular to smooth. Apparently dioicous. Perichaetia terminal, the bracts sheathing at base but otherwise scarcely differentiated or smaller, with rounded tips. Setae elongate; capsules cylindric; operculum erect, long-rostrate; annulus not or variously differentiated; peristome none (or possibly rudimentary).