Plants medium sized, forming rather course mats, dark green to yellowish-green, or golden. Stems spreading or creeping, stems and branches often ascending, sub- to bipinnately branched, flagellate branches present or absent; in cross-section outer 2-3 cell rows small, thick-walled, inner cells large, thin-walled, central strand absent; pseudoparaphyllia foliose; radiculose below. Leaves differentiated between stem and branches, stem leaves spreading to erect, oblong to ovate short-lanceolate, concave, apex short to abruptly long acuminate, base subauriculate or not; margins plane to slightly recurved below, entire to weakly serrulate distally; ecostate or costate, costae short and forked; median laminal cells linear, often vermicular, smooth, thick-walled; basal cells weakly porose or not, reddish-orange; alar region differentiated, cells inflated, 2-several rows, oblong-rectangular; branch leaves erect-spreading, distinctly smaller, oblong-lanceolate to ovate margins entire to distinctly serrate; flagellate branches when present, with reduced appressed scale-like leaves, appressed, oblong-acute, rather coarsely serrate. Sporophytes not observed in the Neotropics.

Plantas medianas, formando matas bastante áspero, verdes oscuras a verde-amarillentas, o doradas. Tallos patentes o reptantes, tallos y ramas a menudo ascendentes, ramificaciones sub- a bipinnadas, ramas flageladas presentes o no; en corte transversal 2-3 filas de células externas pequeñas, con paredes gruesas, células internas grandes, con paredes delgadas, cordón central ausente; pseudoparafilos foliosos; radiculosos abajo. Hojas del tallo y las ramas diferenciadas, hojas del tallo patentes a erectas, oblongas a ovadas corto-lanceoladas, cóncavas, ápices cortos a abruptamente largo acuminados, base subauriculada o no; márgenes planos a escasamente recurvados abajo, enteros a débilmente serrulados distalmente; con o sin costa, costa corta y bifurcada; células mediales de la lámina lineales, a menudo vermiculares, lisas, con paredes gruesas; células basales débilmente porosas o no, naranja-rojizas; región alar diferenciada, células infladas, 2-varias filas, oblongo-rectangulares; hojas de las ramas erecto-extendidas, conspicuamente menores, oblongo-lanceoladas a ovadas, márgenes enteros a conspicuamente serrados; ramas flageladas cuando presentes, con hojas escuamiformes adpresas y reducidas, oblongo-agudas, muy gruesamente serradas. Esporofitos no observados en el Neotrópico.