Plants small, medium-sized, or robust, green, yellowish green, yellowish red, or red, in thin or dense mats. Stems prostrate, irregularly branched, hyaline to yellowish when young, becoming red-dish with age, complanate-foliate or evenly foliate; stems in cross section with 2–5-stratose sclero-dermis, cortical cells enlarged, firm-walled, central strand absent; paraphyllia absent; pseudopara-phyllia absent; axillary hairs 2–4 cells long, basal 1–3 cells short-rectangular, reddish brown, upper 1–2 cells oblong-elongate, hyaline; rhizoids from clusters of initials abaxial to the leaf insertions, smooth, reddish brown, not or sparsely and irregularly branched. Leaves erect, erect-incurved, falcate-secund, or flattened, at times twisted, often rugose when dry, erect-spreading when wet, symmetric or weakly asymmetric, concave, ovate, ovate-lanceolate, oblong-lanceolate to lanceolate, or oblong-ovate, rounded, obtuse, acuminate to long-acuminate; margins denticulate, serrate, or sinuate above, serrulate, subentire, or entire below, plane or narrowly recurved above, at times broadly incurved on one side below; costae double, thin or thick, ending 2/3–3/4 the leaf length, diverging throughout, smooth on back, at times projecting at the tips; cells smooth or prorate, upper cells rhomboidal, long-rectangular to linear-flexuose, firm-walled; basal cells near the insertions oblong-rectangular to long-rectangular, often porose; alar cells undifferentiated. Dioicous. Setae elongate, smooth or smooth below, roughly papillose to tuberculate on capsule necks and upper parts of setae. Capsules inclined, symmetric and cylindric, neck well developed; exothecial cells oblong to rounded-rectangular, in-crassate, more or less collenchymatous; rudimentary 2–6-celled stomata on neck; opercula long-rostrate, conic at base; annuli massive, persistent, of 4–8 rows of yellowish, hexagonal cells; peristome diplolepideous; exostome teeth narrowly triangular, with broad median furrows, densely and closely striate below, coarsely papillose above, ventral surface hyaline to yellowish, smooth to lightly papil-lose; endostome as long as the exostome, yellowish, lightly papillose throughout, basal membranes high, segments broad, strongly keeled, cilia absent or rudimentary. Calyptrae mitrate, irregularly lobed, reddish and lightly roughened, yellowish below.