Stems and branches terete to complanate-foliate, prostrate or erect-ascending, simple or irregu-larly branched; stems in cross section with epidermis of 1 row of small, thick-walled cells, cortical cells enlarged, central strand absent; paraphyllia absent; pseudoparaphyllia absent; rhizoids from small clusters of initials abaxial to the leaf insertions, papillose-roughened, not or sparsely and irreg-ularly branched. Stem and branch leaves similar, pseudodistichous, concave to asymmetrically conduplicate, rounded to shortly auriculate or straight at base, not decurrent; margins serrulate above, entire below or entire throughout; costae short and double, or absent; cells linear-flexuose, smooth, non-porose; alar cells undifferentiated. Asexual reproduction occasionally by uniseriate, rhizoidal gemmae. Dioicous. Setae elongate, smooth. Capsules erect or inclined; exothecial cells not or some-what collenchymatous; stomata phaneroporic, long-pored; opercula conic to conic-rostrate; annulus present; peristome diplolepideous, exostome narrowly triangular, median line faint, densely cross-striate and papillose on lower dorsal surface, papillose above; endostome papillose, basal membranes high, segments narrow, keeled, perforate, cilia 1–3. Calyptrae cucullate.