Leptodontium exasperatum Card., Rev. Bryol. 36: 74. 1909.
Leptodontium viticulosoides (P. Beauv.) Wijk & Marg. var. exasperatum (Card.) Zand., Bryologist 75: 254. 1972. Protologue: Mexico. Etat de Vera-Cruz: Orizaba (F. Müller). Sans indication de localité (Diguet, in herb. Mus. Paris). Lectotype (Zander 1972): Müller, s.n. (NY).
Plants medium to robust, in loose tufts, yellow-green to light green, to 9 cm high. Stems red, erect or laxly ascending, sclerodermis present, moderately to densely radiculose. Leaves moderately spaced, erect at base, spirally twisted to contorted above when dry, squarrose to squarrose-recurved when wet, ovate-lanceolate to long-lanceolate, 3–5 mm long, keeled above, sheathing at base, decurrent; apices acute; margins recurved in lower 1/2–2/3, dentate in upper 1/3, teeth pluripapillose; costa subpercurrent, usually with high papillae on upper dorsal surface; upper leaf cells quadrate to hexagonal, thick-walled, collenchymatous, 7.5–12.5 μm, with long, single, columnar, simple to bifid papillae, 4–10 μm high, centered over the lumina, inner basal cells long-rectangular to elongate, 30–100 x 5–8 μm, pluripapillose, thick-walled, porose, outer basal leaf cells similar to upper leaf cells, alar cells subrectangular to quadrate, thick-walled, porose, yellow-red. Sporophytes not seen from Central America, description based on Peruvian plants (Allen 4327, MO). Dioicous (?). Setae straight to flexuose, 1 per perichaetium, 10–15 mm long, yellow, becoming brownish with age. Capsules erect, narrowly cylindrical, 3–4 mm long, smooth when dry; exothecial cells short-rectangular, firm-walled; stomata at base of urn; opercula erect, conic-rostrate, 1.5 mm long; peristome teeth linear, divided to the base, red-brown, obliquely striate to slightly papillose on outer surface. Spores heterosporous, 5–10 μm and 12.5–15 μm, slightly papillose. Calyptrae cucullate, smooth, 5–6 mm long.