4. Leptodontium viticulosoides (P. Beauv.) Wijk & Margad. 薄齿藓 bo-chi xian
Taxon 9: 51. 1960. Neckera viticulosoides P. Beauv., Prodr. Aethéogam. 78. 1805. Type. Réunion, Bory de St Vincent s.n.
Leptodontium squarrosum (Hook.) Hampe in Lindb., Öfvers. Förh. Kongl. Svenska Vetensk.-Akad. 21: 227. 1864. Didymodon squarrosus Hook., Musci Exot. 2: 150 f. 1–8. 1819.
Leptodontium squarrosum var. abbreviatum (Dixon) P.C. Chen, Hedwigia 80: 320. 1941. Leptodontium viticulosoides var. abbreviatum (Dixon) Wijk & Margad., Taxon 11: 221. 1962. Leptodontium abbreviatum Dixon, Notes Roy. Bot. Gard. Edinburgh 19: 284, f. 4. 1938.
Leptodontium subdenticulatum (Müll. Hal.) Paris, Ind. Bryol. 732. 1894. Leptodontium squarrosum var. subdenticulatum (Müll. Hal.) P.C. Chen, Hedwigia 80: 320. 1941. Trichostomum subdenticulatum Müll. Hal., Syn. Musc. Frond. 2: 626. 1851.
Leptodontium taiwanense Nog., J. Jap. Bot. 20: 144. 1944, syn. nov. Type. China: Taiwan, Mt. Ari-Tataka, 8 Aug. 1932, H. Ozaki s.n. (holotype NICH).
Plants rather large, to 60 mm high, yellowish green to dark brown, in loose tufts. Stems erect or erectopatent, irregularly branched, often with many reddish brown, leafless branches. Leaves appressed to more or less contorted when dry, curved, keeled when moist, ovate-lanceolate, 0.9–1.6 mm × 0.25–0.65 mm, narrowed at the base, gradually acuminate at the apex; margins narrowly revolute, entire below, irregularly serrulate in the upper half; costa slender, ending just below the apex; upper leaf cells rounded quadrate or irregularly oblong-hexagonal, 10–14 µm × 8–12 µm, thick-walled, densely pluripapillose; basal cells larger, rectangular to narrowly rectangular, 24–30 µm × 6–10 µm, hyaline, rarely yellowish brown. Perichaetial leaves similar to stem leaves. Setae straight, 15–20 mm long, yellowish red; capsules ovoid-cylindrical; annuli differentiated, of 5–6 rows of cells; opercula conic-rostrate, with a long oblique beak; peristome teeth linear-lanceolate, short.