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Published In: Journal of the Hattori Botanical Laboratory 39: 519. 1975. (J. Hattori Bot. Lab.) Name publication detail

Project Name Data (Last Modified On 2/15/2011)
Acceptance : Accepted
Project data     (Last Modified On 2/15/2011)

Didymodon icmadophilus is known in Central America from a single collection. The Central American material is a small representation of the species with leaves that are wide-spreading when moist. This collection closely approaches small expressions of D. acutus from Central America, and may in fact represent that species. Didymodon acutus differs from D. icmadophilus in having longer, broader leaves with a costa that is less stoutly excurrent, margins  recurved to near midleaf, and somewhat larger, often unipapillose leaf cells that are more strongly differentiated at the leaf base.

Illustrations: Bartram (1949, Fig. 55 E–G); Lawton (1971, Pl. 38 6–11); Saito (1975, Fig. 56 15–27); Smith (1978, Fig. 117, 6–9); Zander (1981, Pl. 2 11–13); Sharp et al. (1994, Fig. 224); Churchill and Linares (1995, Fig. 153 g–j). Figure 24.
Habitat: On tree trunks; 3300 m.
Distribution in Central America: GUATEMALA. San Marcos: Steyermark 35816 (F, FH).
World Range: Subarctic America, Western and Eastern Canada, Northwestern, North-Central, Southwestern, and South-Central U.S.A;  Mexico; Central America; Western South America; Northern, East, Southwestern, and Southeastern Europe; Siberia, Caucasus, Middle Asia, Mongolia, China, Western and Eastern Asia; Indian Subcontinent.


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Didymodon icmadophilus (Schimp. ex C. Müll.) Saito, J. Hattori Bot. Lab. 39: 519. 1975.

Barbula icmadophila Schimp. ex C. Müll., Syn. Musc. Frond. 1: 614. 1849. Tortula icmadophila (Schimp. ex C. Müll.) Lindb., Öfvers. Förh. Kongl. Svenska Vetensk.-Akad. 21: 249. 1864. Barbula gracilis ssp. icmadophila (Schimp. ex C. Müll.) Amann, Fl. Mouss. Suisse 2: 105. 1919.  Barbula gracilis var. icmadophila (Schimp. ex C. Müll.) Mönk., Laubm. Eur. 289. 1927.  Barbula acuta ssp. icmadophila (Schimp. ex C. Müll.) Podp., Consp. Musc. Eur. 209. 1954. Didymodon rigidulus var. icmadophilus (Schimp. ex C. Müll.) Zand., Cryptog. Bryol. Lichénol. 2: 394. 1981. Protologue: Austria. Ad catarractam “Krimmlfall” Pinzgoviae societate Bryi julacei, ubi legit W. P. Schimper.  

Plants small, dull, yellow-green above, brown below, to 20 mm high. Stems reddish yellow, erect, irregularly branched, sclerodermis and central strand well-developed; rhizoids sparsely distributed at base of stems. Axillary hairs 3–4 cells long, basal cell brown. Leaves 0.8–1.4 mm long, lanceolate to ovate-lanceolate, spreading at base, well- spaced on stems, erect to erect-incurved when dry, wide-spreading when wet, plane along the costa on ventral surface; apices subulate; lamina unistratose, occasionally bistratose in patches on upper margins; margins entire, plane; costa stoutly excurrent, surface ventral cells quadrate, guide cells present, ventral and dorsal stereid bands present below, only dorsal stereids above, ventral and dorsal surface layer not enlarged; upper cells in regular rows, rounded to oblate, 4–6 x 6–8 μm, firm-walled, smooth, basal cells oblate, rounded or short-rectangular, firm-walled, smooth, 4–10 x 6 μm, extreme basal cells enlarged to 22 x 8 μm, alar cells not differentiated. Dioicous. Sporophytes not seen.



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