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Published In: Phytologia 41: 22. 1978. (Phytologia) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 1/27/2013)
Acceptance : Accepted
Project Data     (Last Modified On 10/6/2017)
Plant Category: Mosses
country distribution: Venezuela, Colombia, Ecuador, Peru, Bolivia, NW Argentina
EcoRegions: Páramo, Puna, Open montane, Dry Inter-Andean Valleys
elevation: 500-1000, 1000-1500, 1500-2000, 2000-2500, 2500-3000, 3000-3500, 3500-4000, 4000-4500
substrate: soil, rocks

Notes     (Last Modified On 10/6/2017)
Ecology: Elevation: 2000-3300 m (Venezuela: 2900-3300 m, Colombia: 2000 m, Peru: 2000-3000 m, Bolivia: 2780 m, Argentina: 2090 m). Distribution: United States, Central America, and Andes; also Europe. Previously recognized as a variety of T. australasiae, see comments under that species.
Didymodon umbrosusis characterized by the presence of a stem hyalodermis, long lanceolate to oblong-lanceolate leaves to 3 mm, acute to obtuse apices, entire, plane margins, unistratose lamina or bistratose in distal 1/3 of leaf, subpercurrent to percurrent costa, dorsal stereid band present, ventral stereid band absent, rectangular upper adaxial costal (epidermal) cells, and differentiated marginal base cells in 2-6 rows (narrowly long rectangular and firm-walled). Previously recognized as a variety of T. australasiae, see comments under that species. Two species described from northwest Argentina including Asteriscium flavisetum Herzog based on a collection made by Hosseus from Orán (Province of Salta), and Barbula curvipes Müll. Hal. based on a collection by Lorentz from Siambon (Province of Tucumán), are synonyms.
Both D. australasiae and D. umbrosa (the latter as a variety of the former in some treatments) have been treated as Trichostomopsis. These two taxa can be separated by the following key (Jiménez et al., 2005):
1. Stem hyalodermis absent or occasionally present in patches; leaves oblong-lanceolate to lanceolate, ovate or triangular; basal cells at margin not or weakly differentiated, quadrate to short rectangular … Didymodon australasiae
1. Stem hyalodermis present; leaves long-lanceolate; basal cells at margin distinctly differentiated, long rectangular … Didymodon umbrosa
Didymodon umbrosus se caracteriza por la presencia de hialodermis en el tallo, hojas largo lanceoladas a oblongo-lanceoladas hasta 3 mm, ápices agudos a obtusos, márgenes enteros, planos, lámina uniestratificada o biestratificada en 1/3 distal de la hoja, costa subpercurrente a percurrente, franja dorsal de estereidas presente, franja ventral de estereidas ausente, células superiores adaxiales de la costa (epidérmicas) rectangulares, y células marginales de la base diferenciadas en 2-6 filas (angostamente largo rectangulares y con paredes firmes). Previamente consideradas variedades de T. australasiae, ver comentarios en esta especie. Dos especies descritas del noroeste de Argentina incluyendo Asteriscium flavisetum Herzog basada en una colección realizada por Hosseus en Orán (provincia de Salta), y Barbula curvipes Müll. Hal. basada en una colección realizada por Lorentz en Siambon (provincia de Tucumán), son sinónimos.
Tanto D. australasiae como D. umbrosa (la última es una variedad de la primera según algunos tratamientos) han sido consideradas como Trichostomopsis. Ambos taxa pueden ser separados mediante la siguiente clave (Jiménez et al., 2005):
1. Tallos sin hialodermis u ocasionalmente presente en parches; hojas oblongo-lanceoladas a lanceoladas, ovadas o triangulares; células basales en el margen no o débilmente diferenciadas, cuadradas a corto rectangulares … Didymodon australasiae
1. Tallos con hialodermis; hojas largo-lanceoladas; células basales en el margen conspicuamente diferenciadas, largo rectangulares … Didymodon umbrosa



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