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Published In: Muscologiae Recentiorum Supplementum 4: 73. 1819[1818]. (Muscol. Recent. Suppl.) Name publication detailView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 4/12/2011)
Acceptance : Accepted
Project data     (Last Modified On 4/12/2011)
Discussion:

Campylopus richardii is recognized by its leaves with hyaline hair-tips, short, incrassate, porose leaf cells, and the frequent absence of costal ventral stereid cells. When ventral stereids are absent the ventrally exposed costal guide cells can be mistaken for hyalocysts. Previously excluded from Guatemala (Allen, 1989), the species has since been confirmed for that country. It may be confused with C. pilifer, C. paramoensis, C.valerioi, or the hair-tipped expression of C. savannarum. The first two have ventral hyalocysts in their costae. Campylopus savannarum has non-porose leaf cells and quadrate basal cells. Campylopus valerioi is a larger plant with leaves at base having a short hyaline border of elongate cells.

Illustrations: Bartram (1949, Fig. 22 D–H); Frahm (1978, Pl. 21); Frahm (1979, p. 177); Frahm and Gradstein (1987, Pl. 1 d); Frahm (1991a, Fig. 113).
Habitat: On gravel and soil, road banks and exposed rocks; 900–3447 m.
Distribution in Central America: GUATEMALA. Chimaltenango: Standley 61676 (FH). EL SALVADOR. Santa Ana: Watson 46 (MO). HONDURAS. Comayagua: Allen 11024 (MO, TEFH); Cortés: Allen 14052 (MO, TEFH); Lempira: Allen 12185 (MO, TEFH); Ocotepeque: Allen 14526 (MO, TEFH); Olancho: Allen 12704 (MO, TEFH). COSTA RICA. Cartago: Crosby & Crosby 6134 (MO); Heredia: Crosby 10933b (MO); San José: Crosby 9828 (MO). PANAMA. Bocas del Toro: Allen 5286 (MO); Chiriquí: Allen 9099 (MO); Panamá: Allen 5314 (MO).
World Range: Mexico; Central America; Caribbean, Western and Northern South America, Brazil.

 

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Campylopus richardii Brid., Muscol. Recent. Suppl. 4: 73. 1819. Thysanomitrion richardii (Brid.) Schwaegr., Sp. Musc. Frond. Suppl. 2: 61. 1823. Dicranum richardii (Brid.) C. Müll., Syn. Musc. Frond. 1: 413. 1848. Pilopogon richardii (Brid.) Broth., Nat. Pflanzenfam. 1(3): 336. 1901. Type. Guadeloupe, Parker.

Plants medium to robust, yellowish green at tips, dark brown to black below; stems 3–6 cm long. Leaves on fertile stems appressed below, in comal tufts above, oblong-lanceolate, 3–8 mm long, apex acute, hyaline hair-pointed; margins concave below, tubulose above, entire; costa excurrent, mammillose at back, in cross section with small guide cells, dorsal stereid cells well-developed, ventral stereid cells weakly developed, consisting of only a single layer or absent, if absent guide cells ventrally exposed; upper cells obliquely rhomboidal; basal cells linear-rectangular, all incrassate-pitted; alar cells forming distinct groups of red-yellow cells. Setae 4–8 mm long, red-black, rough above; capsules erect, symmetric, cylindrical-ellipsoid, 1.5–2.0 mm long, scabrous at base, red-black; opercula rostrate, 1 mm long. Calyptrae 2 mm long, ciliate at base.

 

 

 
 
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