Plants general small to somewhat medium sized, dark to yellow-green, in mats, spreading and sometimes subascending, irregularly to irregularly pinnately branched, n cross-section hyalodermis absent, central strand present, pseudoparaphyllia foliose. Leaves erect to somewhat spreading, ovate-lanceolate to ovate, 0.6-2 mm, acuminate to acute, margins weakly denticulate to entire, costa slender to strong, subpercurrent to percurrent, rarely short excurrent, upper median cells oblong-rhombic, relatively short, 2-4:1, walls firm, somewhat thin to thick, basal cells larger, alar region weakly differentiated, cells mostly short or somewhat long rectangular to quadrate above; branch leaves somewhat smaller. Autoicous. Setae 10-25 mm, smooth. Capsules curved and asymmetrioc, to 2.5 mm; peristome double, exostome cross-striolate below, endostome segments ca. equal to exostome teeth.