Plants medium-sized to robust, 3–10 cm high, stipitate-frondose, flattened, forming pale green or yellowish green, lax tufts. Stolons creeping; rhizoids smooth, reddish brown, densely present on stolons and at the base of stipitate stems; leaves appressed, widely spaced, evenly foliate, triangular-lanceolate. Secondary stems erect, stipitate, irregularly bi- or tripinnate, complanate-foliate, at times flagellate-attenuate; secondary stems in cross section with sclerodermis of 3–6 rows of small, thick-walled cells, cortical cells enlarged, firm-walled, central strand absent; paraphyllia absent; pseudopa-raphyllianarrowly foliose to filamentous. Stipe leaves erect-spreading from an appressed base, evenly foliate, triangular to oblong-triangular, 1.5–2.0 mm long, broadly acuminate; margins entire to suben-tire below, serrate above; costae single, ending above midleaf. Stem leaves 2–4 mm long, branch leaves 1.0–1.5 mm long, both flattened, spreading to wide-spreading, smooth when dry or wet, broadly oblong-ovate, oblong-ligulate, obovate, or oblong-spathulate, asymmetric, broadly rounded and coarsely toothed at apex, teeth consisting of 4–8 rows of rhomboid cells, terminal tooth usually longer than the lower teeth, narrowly decurrent, straight at the insertion; margins inflexed on one side below, entire to subentire below; costae slender, often spurred below, tapering and occasionally forked above, reaching 1/2–2/3 the leaf length; cells smooth, apical cells rounded-rhomboid to hex-agonal, or shortly fusiform, 12–20 µm long, firm- to thick-walled, occasionally porose, median cells oblong-rhomboidal, linear-rhomboidal, or long-rectangular, 30–50 µm long, thick-walled, porose, basal juxtacostal cells linear-fusiform, thick- to firm-walled, porose, basal marginal cells rectangular to subquadrate; alar cells rectangular to oblong-rectangular, weakly differentiated, cells across the in-sertions porose. Asexual reproduction by microphyllous branches. Dioicous. Sporophyte unknown in the Neotropics, description from Ninh (1984) and Enroth (1989). Perichaetia to 2 mm long; leaves ovate at base, ligulate to long lanceolate and spreading above, dentate at apex. Setae 1.5–4.5 mm long, smooth throughout or smooth below and mammillose above. Capsules ovoid-cylindrical, 1.5–3.0 mm long; exothecial cells near mouth subquadrate, thick-walled, short-rectangular, thick-walled below; stomata on neck; opercula conic, obliquely short-rostrate; annuli weakly differentiated; peristome diplolepideous; exostome teeth linear-lanceolate, 0.6–0.7 mm long, papillose throughout; endostome basal membranes short, segments similar to exostome teeth in shape and size, cilia not noted. Spores 14–20 µm, papillose. Calyptrae cucullate, sparsely hairy, smooth.