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Didymodon crassicostatus (E.B. Bartram) R.H. Zander Search in The Plant ListSearch in NYBG Virtual HerbariumSearch in Muséum national d'Histoire naturelleSearch in Type Specimen Register of the U.S. National HerbariumSearch in Virtual Herbaria AustriaSearch in JSTOR Plant ScienceSearch in SEINetSearch in African Plants Database at Geneva Botanical GardenAfrican Plants, Senckenberg Photo GallerySearch in Flora do Brasil 2020Search in Reflora - Virtual HerbariumSearch in Living Collections Decrease font Increase font Restore font

Published In: Bulletin of the Buffalo Society of Natural Sciences 32: 162. 1993. (Bull. Buffalo Soc. Nat. Sci.) Name publication detail

Project Name Data (Last Modified On 2/15/2011)
Acceptance : Accepted
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In technical features D. crassicostatus is somewhat like D. acutus or D. laevigatus, and the species has been treated as a synonym of D. laevigatus (Sollman 1994). Didymodon crassicostatus differs from those species in its longer leaves, stouter costae (over 100 μm wide at base), and better developed costal stereid bands. The presence of smooth leaf cells, very broad costae, and exceptionally long leaves distinguish it from all other Central American species of Didymodon. There are only two collections of this species, both grew near a waterfall in the same locality. Plants from the holotype of D. crassicostatus, as noted by Bartram (1949), have the general aspect and fragile leaves of Pseudosymblepharis. All Central American species of Pseudosymblepharis differ from this species in having thinner-walled leaf cells that are papillose above and enlarged at base. Plants in the other collection of this species are more compact, have less fragile leaves, and a more Didymodon-like aspect.

Illustrations: Bartram (1949, Fig. 55 H–K). Figure 21.
Habitat: Moist shaded bluffs by waterfall, in direct spray; 2700 m.
Distribution in Central America: GUATEMALA. San Marcos: Steyermark 35695
World Range: Central America.


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Didymodon crassicostatus (Bartr.) Zand., Bull. Buffalo Soc. Nat. Sci. 32: 162. 1993.

Barbula crassicostata Bartr., Bryologist 49: 114. 1946. Protologue: Guatemala. Dept. San Marcos: along road between San Sebastián at km. 21 and km. 8, 8–18 miles northwest of San Marcos, alt. 2700–3800 m, Steyermark 35695 (F).


Plants robust, dull, green above, brown below, in loose tufts, 30–40 mm high. Stems red erect, sparsely and irregularly branched, sclerodermis and central strand present; rhizoids sparse, at base of stems or scattered throughout. Axillary hairs 4–5 cells long, basal cell brown. Leaves 4–6 mm long, linear-lanceolate to triangular-ovate, spreading at base, circinate-contorted above when dry, spreading to falcate-secund above when wet, plane along the costa on ventral surface; apices subulate; lamina unistratose, fragile; margins entire, recurved below, plane above; costa stout, 100–140 μm wide at base, filling the apex and percurrent to excurrent, ventral surface cells of costa quadrate, guide cells and two stereid bands well-developed, ventral and dorsal surface cells enlarged; upper and basal cells weakly differentiated, upper cells in regular rows, rounded, 8–10 x 8 μm, firm-walled, smooth, basal cells quadrate, rounded or short-rectangular, firm-walled, smooth, 10–30 x 10 μm, alar cells not differentiated. Sporophytes not known.


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