Macromitrium crosbyorum Allen & Vitt in Allen, Novon 8: 113. 1998.
Protologue: Costa Rica. San José: along Inter American Highway, ca. 10 km NW of summit at La Ascensión, 9°37'N, 83°48'W, Crosby & Crosby 6089 (ALTA, CR, MO, NY, US).
Plants large, greenish red to yellowish red. Stems creeping to 7 cm, branches 2–3 cm long, reddish tomentose below. Leaves keeled, erect below, flexuose to spirally contorted and undulate above when dry, erect‑patent when wet, (3–)4–6 mm long, 1 mm wide, lanceolate; apices acuminate; margins undulate, serrate above, frequently serrulate to near the base, recurved below, erect to plane above, enlarged basal teeth at leaf insertion absent; costae percurrent; upper interior cells 8–20 μm, rounded and collenchymatous, isodiametric to rhombic, stoutly unipapillose to mammillose, upper marginal cells narrow and elongate forming a more or less distinct border, basal cells long-rectangular, incrassate and porose, densely tuberculate, 26–44 μm long. Dioicous. Setae 7–10 mm long, smooth. Capsules 1.5–2.0 mm long, ovoid to cylindrical, plicate; opercula rostrate, 1–1.5 mm long; annuli non‑revoluble, with fragments adhering to capsule mouth; exostome teeth truncate, 320–424 μm high, yellow, densely papillose‑striate, united and forming a membrane, more or less reflexed at tips, splitting into eight pairs of teeth with age, endostome hyaline, lightly papillose, basal membrane 80–90 μm high, segments 60–80 μm high. Spores anisosporous, 14–20 μm, smooth to lightly papillose and 30–48(–54) μm, densely papillose. Calyptrae mitrate, deeply laciniate, naked, 4–5 mm long.