Pseudosymblepharis richardsii (Bartr.) Allen, comb. nov.

Tortella richardsii Bartr., Bryologist 49: 112. 1946. Protologue: Guatemala. Dept. Alta Verapaz: vicinity of Cubilgüitz, 1½ –2 miles south of Cubilgüitz, alt. 300–350 m, Steyermark 44389 (F). 

Plants medium-sized, dull yellow-green to dark-green, in tufts to or cushions, to 25 mm high. Stems red, erect, sparsely and irregularly branched, hyalodermis present, central strand weakly developed; rhizoids moderately developed below, reddish brown, roughened-papillose. Leaves 5–8 mm long, erect or weakly clasping at base, incurved, spirally twisted and contorted, tubulose above when dry, spreading when wet; apices acute to acuminate, sharply mucronate; laminae fragile and often broken or eroded above; margins entire and plane; costa short excurrent, guide cells and two stereid bands well-developed, ventral surface layer enlarged; upper cells sub-quadrate to short rectangular, 16–10 x 8–10 μm, firm-walled, pluripapillose, those near the costa extending downward considerably more than those toward the margins, basal cells rectangular, thin-walled, enlarged, bulging, hyaline, smooth, rarely a few cells near costa at the junction of the sheath and limb thick-walled and porose, 85–30 m x 10–25 μm, outer basal cells running up the margins and forming a strong v-shaped pattern 1/2–2/3 the leaf length. Setae 1 per perichaetium, elongate, smooth, spirally twisted, 8–10 mm long, yellow, becoming red with age. Capsules narrowly cylindrical, 1.5–2 mm long, smooth to lightly wrinkled when dry; stomata in neck; opercula erect-rostrate, 1 mm long; annuli well-developed; peristome teeth 16, short, erect to weakly twisted, papillose. Spores 13–15 μm. Calyptrae smooth, 4 mm long.