Accepted By:
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Lee, G. E., S. R. Gradstein, E. Pesiu & N. Norhazrina Binti Nik Mohamed Kamil. 2022. An updated checklist of liverworts and hornworts of Malaysia. PhytoKeys 199: 29-111.
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Wang, J., R.-L. Zhu & S. R. Gradstein. 2016. Taxonomic revision of Lejeuneaceae subfamily Ptychanthoideae (Marchantiophyta) in China. Bryophyt. Biblioth. 65: 4–141.
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