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(Last Modified On 5/20/2013)
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(Last Modified On 5/20/2013)
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Species
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Adenocalymma arthropetiolatum A. Gentry
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Note
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TYPE: Panama, Canal Zone, Pipeline Road, Gentry 2460 (MO).
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Description
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Liana to 3 cm in diameter, the bark smooth, greenish, with inconspicuous lenticels; branchlets subterete to subtetragonal, glabrous to subscabrous, drying tan to greenish, pseudo-stipules small, 4-scaled, usually narrow. Leaves 3-folioate or 2-foliolate with a tendril or tendril scar; leaflets drying olive, narrowly ovate to almost linear, acute, basally truncate, 5.2-17 cm long and 1.9-7.5 cm wide, ? chartaceous, secondary veins 5-7 on each side, above scabrous along the midvein, beneath almost glabrous or puberulous along main veins and sometimes sparsely
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Habit
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Liana
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Description
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on the surface, with q. few scattered plate-shaped glands; tendril simple, to 14 cm long; petiolules 0.5-4.1 cm long, the petiole 1.1-6.4 cm long, puberulous, woody proximally and conspicuously jointed with the petiolules. Inflorescence an axillary or terminal raceme, a subtending glandular, deciduous bract enclosing each bud, the rachis and pedicels puberulous. Flowers with a faint musky odor, the calyx cupular, 5-toothed but more or less bilabiate, 5-15 cm long and 4-10 mm wide, puberulous with simple and thick-stellate trichomes, usually with conspicuous submarginal sunken cupular glands; corolla uniformly bright yellow, often a slightly deeper shade within, campanulate, 4.2-8.2 cm long and 1.1-2.1 cm wide at the mouth, the base cylindrical, 2.3-2.6 cm long and 0.6-0.7 cm wide, the tube 2.5-5.6 cm long, the lobes often folded or rolled, the upper shorter two lobes 1.2-1.7 cm long, extending horizontally in front of the tube, the longer three lobes 1.8-2.3 cm long, variously reflexed, outside and inside the lobes puberulent with simple and thick-stellate trichomes, the tube glabrous inside except for slender 2-3-celled gland-tipped trichomes at and just below the level of stamen insertion, the base of the lobes with plate-shaped glands outside; stamens slightly exserted, the anthers held underneath the hood formed by 2 projecting upper corolla lobes, the anther thecae more or less divergent, 5-6 mm long, the filaments more or less equal, the anterior pair 3.5-3.9 cm long, the posterior pair 3.3-4.0 cm long, the staminode 12-21 mm long, inserted 2.1-2.6 mm from base of corolla tube, pistil 5.0-5.9 cm long, the ovary narrowly cylindrical, 4-5 mm long and 1.5-2.0 mm wide, puberulent, the ovules 2-seriate; disc pulvinar-cupulate, 1.5-2.5 mm long, 4-6 mm wide. Capsule unknown.
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Distribution
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Apparently restricted to stream banks and lake shores of the tropical moist forest and premontane wet forest of central Panama,
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Native
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Panama
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Note
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this plant flowers during the late wet season in October and November and is thus temporally isolated from its congener A. apurense. The smooth greenish bark and wiry, strongly bent petiole and petiolule joints are distinctive. Narrowly ovate leaflets which are slightly 3-nerved at the base also aid in distinguishing it from A. apurense. This species is most closely related to A. magdalenense Dugand of the Mag- dalena Valley of Colombia, which is vegetatively distinguishable mainly by its much larger, glandular, subfoliaceous pseudostipules. The corolla of A. mag- dalenense, however, is evenly and narrowly tubular-infundibuliform with strongly exserted anthers as opposed to A. arthropetiolatum which has its corolla abruptly tubular-campanulate above a narrowly tubular base and subexserted anthers held under a hood formed by the forward-projecting upper corolla lobes. Adenocalymma arthropetiolatum is unusual in the family and especially in the Bignonieae in its restricted geographical range. Since all collections of this species have come from along rivers or lakes, the seeds, when discovered, may prove to be water dispersed. Probably because of this it is restricted to the drainages of the Rios Bayano and Chagres plus the adjacent Rios Pasiga and Maestro.
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Specimen
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CANAL ZONE: Pipeline Road, Gentry 1435, 2460 (both MO); Barro Colorado Island, Standley 40912 (US). PANAMA: Rio Aguacate near El Llano, Duke 5828 (MO, SCZ). Near mouth of Rio Pasiga, Gentry 2205A (MO). Rio Maestro, Gentry 2240 (MO). Rio Piragua This content downloaded from 192.104.39.2 on Tue, 14 May 2013 16:10:35 PMAll use subject to JSTOR Terms and Conditions1973] GENTRY--FLORA OF PANAMA (Family 172. Bignoniaceae) 791 (Tabardi), Gentry 2519 (MO). Between Rios Espave and Agua Clara, Gentry 3772 (MO). Near Madden Lake, Gentry 5011 (MO).
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