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Published In: Species Plantarum. Editio quarta 3(1): 400. 1800. (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
General/Distribution: A genus of c. 100 species, distributed in tropical and subtropical regions of Africa and Asia; represented in Pakistan by 7 species.


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Prostrate to erect, mostly pubescent herbs or subshrubs with nodally thickened stems, woody rootstock present or absent. Leaves cauline, sessile or petiolate, narrowed at both ends, entire or crenate-serrate. Flowers small, sessile to subsessile, in terminal or axillary, dense, spinescent or sericeous, condensed spikes or sometimes in globose, capitate, ± radical heads; bracts conspicuous, lanceolate or elliptic, sharp acute or spinescent; bracteoles 2 or absent, smaller and similar to bracts. Calyx deeply 5-lobed, lobes unequal, anterior lobes mostly basally connate making the calyx apparently 4-lobed, imbricate, mucronate. Corolla sericeous outside, tube uniformly cylindrical below, constricted in the middle, suddenly swollen above, as long as or scarcely baler than the patent limb, 2-lipped, upper lip 2-fid, erect to patent, lower lip 3-lobed, lobes imbricate. Stamens 4, didynamous, inserted at the base or throat of corolla tube, included or exserted, all fertile, anthers oblong, minutely ciliate, dithecous, thecae equal or unequally placed, basally acute. Ovary globose, bilocular, each locule 2(-1)-ovuled; style filiform with minutely 2-fid stigma. Capsule ellipsoid or oblong, 2-4-seeded, apically hairy. Seeds flattened, ovate-oblong or ± orbicular. hairy.

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1 Leaves more than 1 cm broad (2)
+ Leaves less than 1 cm broad (5)
2 (1) Calyx 5-lobed. Leaves short to long petioled, broadly lanceolate-oblong to ovate-elliptic (3)
+ Calyx 4-lobed. Leaves almost sessile, linear to narrowly lanceolate 1 Lepidagathis rigida
3 (2) Leaves, bracts and bracteoles not cuspidate. Seeds 1-1.5 mm long (4)
+ Upper leaves, bracts and bracteoles cuspidate or spine-tipped. Seeds c. 2.5 mm long 2 Lepidagathis cuspidata
4 (3) A prostrate herb with softly pubescent twigs. Leaf Margins crenate 4 Lepidagathis fasciculata
+ A suberect herb with almost glabrous twigs. Leaf margins entire 3 Lepidagathis incurva
5 (1) Capsule 2-seeded (6)
+ Capsule 4-seeded (7)
6 (5) Flowers light pink. Bracts and bracteoles minutely puberulous, less than 1 cm long 5 Lepidagathis trinervis
+ Flowers yellow. Bracts and bracteoles silky pubescent, 1 cm or more long 6 Lepidagathis lutea
7 (5) Calyx 5-lobed. Leaves distinctly petiolate (8)
+ Calyx 4-lobed. Leaves almost sessile 1 Lepidagathis rigida
8 (7) Leaves dimorphic, those on main stem 5-10 (-12) cm long, upper leaves cuspidate, glandular-pubescent 2 Lepidagathis cuspidata
+ Leaves not dimorphic, not more than 2 cm long, cuspidate, almost glabrous 7 Lepidagathis calycina
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