Strophostyles Elliott
(Riley-Hulting et al., 2004)
Plants annual or
perennial herbs, from short taproots, sometimes climbing on other vegetation. Stems trailing or
climbing, twining, branched, sometimes rooting at the nodes, unarmed, moderately
to densely pubescent with short, downward-curved to downward-angled and
sometimes also more or less spreading hairs, these slender to somewhat tapered
from the base, but lacking a bulbous base. Leaves sometimes
drooping and folding at night, pinnately trifoliate,
mostly long-petiolate, the petiole hairy.
Stipules 2–4(–5) mm long, narrowly ovate-triangular, herbaceous to papery,
appearing basally attached and somewhat spreading, mostly persistent, strongly
parallel-veined; stipels 0.5–1.0(–1.5) mm long. Leaflets lanceolate to ovate, the lateral
ones sometimes appearing asymmetrically rhombic or pear-shaped, entire or with
a basal lobe. Inflorescences very short axillary racemes (rarely only a
solitary flower), appearing as small headlike or
umbellate clusters at the tip of a stalk that usually is much longer than the
subtending leaf, the very short flower stalks with small bracts at the base,
each flower also closely subtended by 2 small bractlets.
Calyces with the tube bell-shaped, 2-lipped, the upper 2 lobes fused with only
a shallow notch at the tip, the lower 3 lobes triangular, unequal, the
lowermost lobe somewhat narrower and longer than the others, often as long as
or longer than the tube. Corollas papilionaceous,
pink to lavender (the keel sometimes darker purple), rarely white, often fading
to green or yellow, the petals cordate or abruptly
tapered to an often relatively broad stalklike base,
the banner broadly ovate to nearly circular, rounded to shallowly notched at
the tip, abruptly arched upward above the midpoint but generally appearing
hooded over other petals, the wings oblong, shorter than the keel, the keel
widest at the midpoint, abruptly constricted above the midpoint into a blunt,
upward-arched beaklike tip that is slightly to strongly twisted to the side.
Stamens 10, 9 of the filaments fused to above the midpoint and 1 filament more
or less free, the anthers small, attached at or near the base, all similar in
size. Ovary appearing arched, sessile, the style curved,
jointed above the midpoint, bearded on upper side, the stigma terminal and capitate. Fruits legumes, linear, not or only
slightly flattened, sessile, short-tapered on the lower side to a short slender
beak, the margins otherwise parallel, glabrous or appressed-hairy,
papery in texture, dehiscent, the valves becoming
spirally twisted during dehiscence, 3–10-seeded. Seeds subcylindric,
rectangular in outline, slightly flattened, the surface brown, often mottled
with darker brown or purplish brown, smooth or minutely pubescent with white-woolly
patches of hairs. Three species, North America, mostly in the
U.S.
Strophostyles is very similar and probably closely
related to Phaseolus,
differing mainly in the twisted (vs. coiled) nearly vertical keel, lack of
hooked hairs, short flower stalks, and somewhat swollen nodes of the
inflorescence. Although palatable to deer and livestock, Strophostyles species generally
are not abundant enough to be considered important forage plants. The genus
sometimes is planted for erosion control.