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(Last Modified On 6/5/2013)
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(Last Modified On 6/5/2013)
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Genus
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Hebeclinium DC.
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PlaceOfPublication
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Prodr. 5: 136. 1836.
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Note
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TYPE: Eupatorium macrophyllum L.
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Description
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Erect large herbs or subshrubs, sparingly branched. Leaves always opposite; blades broadly ovate to deltoid, usually serrate; petioles rather long. Inflorescence a corymbose panicle or lax cyme. Heads discoid with 20-80 florets; involucre of 25-40 bracts, in 3-5 series, the outer bracts persistent; receptacle hemispherical, often with center broken off, glabrous to densely hirsute, sclerified throughout (parenchymatous internally in one species); corolla narrowly tubular, 5-lobed, the outer surface of corolla glabrous below, the lobes usually longer than wide, usually with prominent multicellular uniseriate hairs and a few glands, the inner surface of some species with numerous hairs, the corolla cells narrow with sinuous walls; anther collar often slender, with many quadrate cells in lower part, the cell
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Habit
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herbs or subshrubs
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Description
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walls rather thin with indistinct annular thickenings, the anther appendage ovate, longer than wide; style base not enlarged, glabrous, the style appendages narrow throughout, slightly mamillose. Achenes prismatic, often slightly curved, 4-5- ribbed, bristles sometimes present; carpopodium scarcely distinct, only a few rows of short cells at lower margin, the area of longer upper cells merging with sides of achene and extending up ribs; pappus of ca. 30-40 scabrous bristles, the apical cells acute.
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Distribution
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mainly in northern South America.
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Note
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Three species have been collected in Panama. Members of the genus are distinguished primarily by the receptacle but the very slender style appendage is also a notable feature. Hebeclinium has usually been recognized at the sectional level for members of "Eupatorium" having hairs on the receptacle. The concept contained many but not all members of the related generic series, Hebeclinrium, Bartlettina and Decachaeta, where there seems to be a tendency toward pubescent receptacles. Section Hebeclinium suffered most by the inclusion of extraneous elements such as Polyanthina which is related to Ayapana, and Urolepis which is related to Gyptis.
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Reference
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King, R. M. & H. Robinson. 1969. Studies in the Compositae-Eupatorieae, IX. A review of the genus Eupatorium section Hebeclinium in Colombia. Sida 3: 321-326. King & Robinson. 1971. Studies in the Eupatorieae (Asteraceae) XXXVII. The genus Hebeclinium. Phytologia 21: 298-301.
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Key
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a. Heads with 50-80 florets; inflorescence a corymbose-panicle; base of leaf blade usually truncate to cordate ...... 2. H. macrophyllum aa. Heads with 20-40 florets; inflorescence a lax cyme; base of leaf blade cuneate. b. Heads with 20-25 florets; leaf margins doubly serrate ...... 3. H. reedii bb. Heads with ca. 40 florets; leaf margins crenulate ...... 1. H. costaricense
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