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Published In: Species Plantarum 2: 829. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 5/16/2022)
Acceptance : Accepted
Project Data     (Last Modified On 10/24/2020)
Contributor Text: Abdul Ghafoor
Contributor Institution: Centre for Plant Conservation, University of Karachi – PAKISTAN

 

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Annual or perennial, monocarpic herbs or subshrubs, leaves coriaceous, dentate to pinnatisect, marginally spinosely lobed or toothed, in basal rosettes or alternate on shoots, rarely unarmed. Capitula  homogamous, discoid or rarely heterogamous and radiate, solitary, sessile, subtended by foliaceous pectinate bracts. Involucre ovoid or campanulate. Phyllaries spiny, rarely entire, imbricate, outer leaf-like, pectinate, bipinnatisect, inner ones linear-lanceolate, usually with brown, wide scarious appendage. Receptacle flat, beset with hyaline, large, laciniate-fimbriate, free, rigid scales. Florets pink, purple  or yellow, outer ligulate, female, rarely with staminodes or bisexual; inner tubular, bisexual, 5-fid. Anthers basally caudate-sagittate with fimbriate tails, on free filaments. Styles short with erect, stigmatic branches. Cypselas terete to obconical, densely silky hairy, apically without elevated rim, hilum basal. Pappus 1-3-seriate of plumose scales, connate at the base.

A genus of 30 species, distributed in open dry habitats in Canary Islands, Mediterranean region eastwards to Afghanistan and Pakistan. Represented in Pakistan by 4 species.

 

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1.

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Capitula radiate with outer yellow ligulate florets. Outer phyllaries akin to cauline leaves.

 

 

 1. A. carduus

 

Capitula discoid with all florets pink or purple and tubular. Outer phyllaries not akin to cauline leaves.

 

 

2

 

2.

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Annual. Outermost phyllaries finely deeply bipinnatisect,  apically incurved.

 

 

2. A. cancellata

 

Perennial or biennial. Outermost phyllaries not bipinnatisect, apically not incurved                .

 

 

 

3

3.

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Leaves with 2-4 stiffly spiny lobes. Capitula 8 – 12(-15) mm across. Pappus half as long as cypselas.

 

 

3. A. arabica

 

Leaves with more than 4 spiny lobes. Capitula 3 – 5 cm across. Pappus almost as long  as cypselas.

 

4. A. cuneata

 
 
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