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Project Name Data (Last Modified On 5/29/2013)

Flora Data (Last Modified On 5/29/2013)
Genus Voyria Aubl.
PlaceOfPublication Hist. P1. Guiane Fr. 208. 1775.
Synonym Biglandularia Karsten, Linnaea 28: 416. 1856. Leiphaimos Auct. non Schlecht. & Cham., Linnaea 6: 387. 1831.
Description Herbs, saprophytic, devoid of chlorophyll, perennial; the stems usually erect, simple to sometimes branched. Leaves small, scale-like, opposite, decussate, sessile, the opposite pairs usually ? connate at the base. Inflorescences terminal and dichasial, or flowers mostly terminal and solitary. Flowers 4-6-merous; calyx tubular to campanulate, sometimes hyaline, occasionally provided within and at the base with a verticil of small scales, persistent; corolla variously colored, much longer than the calyx, marcescent, the tube elongated, the lobes contorted, spreading; stamens included, the filaments long or the anthers sessile, inserted below the throat of the corolla tube or on the corolla tube at various
Habit Herbs
Description heights, the anthers introrse, coherent or not, the thecae appendaged or not at the base; ovary sessile or shortly stipitate, eglandular or with 2 opposite glands at the base, the latter sometimes stipitate, I-locular, the 2 parietal placentae some- times protruding, style filiform, stigma usually simple, capitate or infundibuli- form, rarely 2-lobed. Capsules surrounded by the persistent calyx and marcescent corolla, ellipsoid to oblong to linear, septicidally 2-valvate or only medially septi- cidal, the valves remaining united at the base and apex; seeds numerous, minute, ovoid and wingless or narrowly fusiform, i.e. provided with 2 long hair-like wings. Pollen grains in monads, bilateral, heteropolar, reniform, semi-ovidal to unsym- metrically biconvex, or radially symmetrical, isopolar, oblate to spheroidal, or irregular in shape, the outline in lateral view convexo-concave to convexo-plane to depressed ovate, or ? circular to irregular, in polar view circular to oval, or irregular, 7-12 x 11-21 x 9-14 microns, or 10-11 x 12-15 microns, or 14-15 J, in diameter, 1-2 porate (one pore often smaller), the pores 1-3 microns in diameter, the pore margin re- inforced by a thickening (annulus) 0.5-1 microns wide, 1-2 microns high, consisting of sexine and/or nexine, the annulus occasionally absent; exine 0.5-1 ,t thick, at the aper- tures up to 2 microns thick, the stratification obscure, non-baculate; sexine smooth or scabrous.
Distribution A tropical genus of about 16 species, mainly in Central and South America and the West Indies, with 1 species in tropical Africa; 9 species reported from Panama.
Note Growing upon decayed leaves on forest floors.
Reference Raynal, A. IRtude critique des genres Voyria et Leiphaimos (Gentianaceae) et revision des Voyria d'Afrique. Adansonia, ser. 2. 7: 53-71. 1967. Robyns, A. Notes on some American species of Voyria (Gentianaceae). Ann. Missouri Bot. Gard. 55: 398-399. 1969. Standley, P. C. The Panamanian species of Leiphaimos. In "Studies of trop- ical American phanerogams-No. 3." Contr. U. S. Natl. Herb. 20: 194-200. 1919. Williams, L. 0. Voyria and Leiphaimos. In "Tropical American plants, IX." Fieldiana: Bot. 31: 411-415. 1968.
Key a. Inflorescences dichasial. b. Herb entirely white; flowers sessile or subsessile; opposite scales connate at the base only; corolla 10-11 mm long, the tube constricted ca. 2 mm below the apex, glabrous within ...... 1. V. alba bb. Herb with orange to purplish-yellow stems; pedicels 2-8 mm long; opposite cauline scales connate for ca. 1/6-/2 of their length; corolla orange or yellow, 15-20 mm long, the tube cylindric, scarcely dilated at the throat, villosulous towards the apex within ...... 2. V. aturantiaca aa. Flowers solitary. c. Calyx without scales at the base within; ovary eglandular. d. Anthers sessile or filaments short, not exceeding 1 mm. e. Anthers exappendiculate. f. Opposite cauline scales connate for ca. 1/5-1/3 of their length or less; cauline scales narrowly ovate, acute apically; calyx lobes narrowly ovate- triangular, acute to acuminate apically; corolla lobes acuminate apically; anthers sessile; ovary substipitate, narrowly ellipsoid ......  3. V. aphylla ff. Opposite cauline scales connate for ca. 1/2-2/3 of their length; cauline scales broadly to very broadly ovate, obtuse to rounded apically; calyx lobes broadly ovate, obtuse apically; corolla lobes obtuse apically; anthers with filaments 0.5-1 mm long; ovary sessile, oblong, apically truncate ...... 4. V. truncata ee. Anthers prolonged basally into hairy appendages ca. 1.5 mm long ...... 5. V. stellata dd. Anthers with the filaments 5-15 mm long. g. Corolla to 2.5 cm long; filaments 5-7 mm long, inserted ca. 1/2 or ca. 2/3 above the base of the corolla tube ...... 6. V. pittieri gg. Corolla to 3.5 cm long; filaments ca. 15 mm long, inserted ca. 1 cm above the base of the corolla tube ...... 7. V. pulcherrima cc. Calyx provided within and at the base with a verticil of emarginate scales; ovary with 2 opposite glands. h. Corolla pale blue; ovary very shortly stipitate, with 2 appressed stipitate glands at the base, the basal part of the stipe adnate to the ovary, the upper part ca. 1-2 mm long and free; stigma capitate ...... 8. V. tenella hh. Corolla orange; ovary sessile, with 2 narrowly elliptic glands ca. 0.5 mm above the base; stigma 2-lobed ...... 9. V. bilobata
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