1. Heuchera L. (alum root)
(Wells, 1984; Kallhoff and Yatskievych, 2001)
Plants with
short, stout rhizomes. Leaves all basal (rarely 1 or a few highly reduced
bracteal leaves alternate on the inflorescence stalks), short- to more commonly
long-petiolate. Stipules scalelike, inconspicuous, fused to the petiole base to
above the midpoint, the margins fringed, persistent on the rhizome after the
leaves die back. Leaf blades mostly about as long as wide, circular to broadly
ovate or kidney-shaped, the base cordate, the tip rounded or narrowed to a
blunt or sharp point, the margins with 3–7 shallow lobes, also finely to
coarsely scalloped or toothed and with short spreading hairs, palmately veined
with usually 7 primary veins, the upper surface glabrous to hairy and/or
glandular, green, sometimes with lighter mottling, the undersurface glabrous to
hairy and/or glandular, usually grayish green or reddish- or purplish-tinged.
Inflorescences panicles with usually numerous flowers, usually long-stalked,
glabrous or more commonly hairy and/or glandular, with small scalelike or
leaflike bracts at the branch points. Flowers somewhat zygomorphic or less
commonly appearing actinomorphic (often appearing more zygomorphic upon
pressing), each subtended by a small, linear bract at the base of the flower
stalk. Hypanthium obconic to bell-shaped, fused to half or more the length of
the ovary. Sepals oblong to triangular-oblong, rounded at the tip. Petals
glabrous to minutely hairy, the margins entire or finely toothed, green, white,
or pink. Stamens 5, barely included to long-exserted from the calyx, the
anthers small, attached toward their midpoints, white, pink, or orange. Ovary
1-locular, the placentation parietal. Styles tapered, persistent and often
arched or spreading at fruiting, the stigmas more or less capitate. Fruits
ovoid, 2-beaked, dehiscing longitudinally from between the beaks. Seeds
numerous, variously shaped, usually somewhat asymmetrical in outline, smooth or
with fine tubercles or spines, dark brown to nearly black. About 35 species,
North America.
Several species
of Heuchera are cultivated to some extent as ornamentals in wildflower
gardens and rock gardens, including all of the species occurring in Missouri.
However, the plants most common in horticulture are the coralbells, with red
flowers, representing H. sanguinea Engelm. (native to the southwestern
United States and adjacent Mexico) and its assortment of cultivars and hybrids.
Heuchera species also have a long history of medicinal use, taken both
internally for various antidiarrheal, astringent, and analgesic properties, and
externally as a dermatological and antirheumatic aid and a styptic (Spongberg,
1972; Moerman, 1998).