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!Cardamine jamesonii Hook.

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Cardamine jamesonii Hooker, London J. Bot. 6: 293. 1847. TYPE: Ecuador, Cordillera of Pilaro, 15,000 ft, Apr 1834, W. Jameson 88 (lectotype, here designated, K!).

Cardamine punicea Turczaninow, Bull. Soc. Imp. Naturalistes Mocou 27(2, 4): 295. 1854 (1855); C. johnstonii Oliver prol. punicea (Turczaninow) O. E. Schulz, Bot. Jahrb. Syst. 32: 420. 1903. TYPE: Venezuela, Merida, Culata, 9-10,000 ft, Apr 1847, Funck & Schlim 1542 (holotype, KW; isotypes, G!, P!).

Cardamine nevadensis Turczaninow, Bull. Soc. Imp. Naturalistes Mocou 27(2, 4): 295. 1854; C. jamesonii Hooker var. nevadensis (Turczaninow) O. E. Schulz, Bot. Jahrb. Syst. 32: 422. 1903. TYPE: Venezuela, Merida, Sierra Nevada, 9,000 ft, Jun 1847, Funck & Schlim 1554 (holotype, KW?; isotypes, G!, 2P!).

Cardamine pulchra Linden & Planchon, Trois. Voy. Linden, Bot., Pl. Columb. 1: 12. 1863. TYPE: Venezuela, Merida, Culata, 9-10,000 ft, Apr 1847, Funck & Schlim 1542 (holotype, P!; isotypes, G!, KW).

Cardamine tolimensis Planchon & Linden in Triana & Linden, Ann. Sci. Nat. Bot. ser. 4, 17: 59. 1862. TYPE: Colombia, Pic de Tolima, près de Boqueron, 3300-3900 m, Jun 1843, Linden 924 (922 in print) (lectotype, here designated, P!).

Cardamine jamesonii var. goudotii Planchon & Linden in Triana & Linden, Ann. Sci. Nat. Bot. ser. 4, 17: 59. 1862. TYPE: Colombia, Mariquita: Toluma, Cuchilla de la Divisadera, 1844, J. Goudot (lectotype, here designated, P!).

Cardamine jamesonii prol. pulcherrima O. E. Schulz, Bot. Jahrb. Syst. 32: 422. 1903. TYPE: Colombia, Cauca: Páramo de Guanacas, 2800-3300 m, F. C. Lehmann 5605 (holotype, B!; isotypes, G!, K!).

Cardamine johnstonii var. superba O. E. Schulz, Bot. Jahrb. Syst. 32: 421. 1903. TYPE: Colombia, Cauca: Páramo de Moras, 3300 m, F. C. Lehmann 2669 (holotype, G-BOIS).

Cardamine rhizomata Rollins, J. Arnold Arbor. 21: 392. 1940. TYPE: Ecuador, Carchi, Nudo de Boliche Voladero, 10 Jun 1939, C. W. Penland & R. H. Summers 870 (holotype, GH!; isotype, COCO).

Herbs, perennial. Rhizomes slender, not scaly, with long internodes and nontuberous nodes. Stems (1.5-)2.8-6.5(-8) dm, erect or decumbent, simple or branched above, glabrous or rarely pubescent. Cauline leaves 6-20; lower and middle leaves (4-)7-14(-18) cm, 7-11(-13)-foliolate, rarely basal ones 5-foliolate; petiole (1.5-)2.5-8(-10) cm, sometimes longer on rhizomal leaves; lateral leaflet (0.5-)1.5-3(-4.5) ´ (0.3-)1-2.2(-3) cm, broadly ovate to oblong-ovate or oblong-lanceolate, glabrous or sparsely to densely pubescent, base strongly oblique and subcordate to cuneate, margin crenate, incised-crenate, or rarely serrate, ciliolate with trichomes to 0.5 mm, teeth mucronate, apex acute; petiolule 0.2-1(-2) cm; terminal leaflet about same size as lateral; uppermost leaves gradually smaller, less divided, and often narrower. Racemes bracteate basally or very rarely only lowermost few flowers bracteate; bracts simple or compound, gradually reduced in size upward; fruiting pedicels (1-)1.5-3 cm, divaricate-ascending, straight, stout. Sepals dark purple, broadly oblong, 3-5 ´ 2-2.5 mm, glabrous; petals purple, violet, red, or rarely pale lavender, broadly obovate, (8-)10-14 ´ 5-8 mm, base broadly cuneate, apex rounded to emarginated; filaments 4-7 mm; anthers oblong, 1-1.5 mm; ovules 16-28 per ovary. Fruits 3-5.5(-7) cm ´ 1.5-2 mm; style stout, 3-8(-10) mm. Seeds oblong,1.5-3 ´ 1-1.2 mm.

Flowering: Sep-Jul.

Habitat: wet trails in forests, páramo, moist banks of quebrada, heath and chapparal vegetationm mixed shrubby paramo, dwarf woodland.

Elevation: 2500-3850 m.

Distribtution: Colombia (Cauca, Mariquita, Nariño, Putumayo, Quindio, Tolima,Valle), Ecuador (Azuay, Carchi, Chimborazo, Cuenca, Imbabura, Loja, Morona-Santiago, Napo, Pichincha,Tungurahua), Vezuela (Merida, Trujillo).