Cardamine lyrata Bunge, Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 6, Sci. Math. 2: 29. 1833.

Cardamine argyi H. Léveillé, Mem. Real Acad. Cienc. Art., Barcelona 12: 547. 1916. TYPE. China, Jiangsu, without locality and collection date, d’Argy s.n. (holotype, E).

            Herbs, perennial, glabrous throughout. Rhizomes short, thick, not scaly; stolons from rhizome or lower to middle stem nodes to 80 cm. Stems (2–)3–7(–8) dm tall erect, simple, striate angled, 8–15-leaved. Leaves on stolons simple or rarely with a basal pair of auriclelike leaflets; petiole 3–12 mm; leaf blade suborbicular, cordate, or reniform, (0.7–)1–2.2(–3) ´ 0.7–2(–2.3) cm, margin repand or entire, apex rounded; cauline leaves sessile, middle ones (1.5–)3–5(–7) cm; terminal leaflet orbicular, ovate, or rarely oblong, 1–3 ´ (0.8–)1–3 cm, with a petiolule (0.3–)0.5–1.5(–2) cm, base cordate, subtruncate, or obtuse, margin repand or subentire, apex rounded; lateral leaflets 2–5 pairs, much smaller than terminal one, ovate, suborbicular, or oblong-ovate; proximal pair of lateral leaflets auriclelike, attached at or just above node, often giving appearance of amplexicaul leaf base. Racemes ebracteate; fruiting pedicels (0.7–)1–2(–2.5) cm, slender, divaricate, usually straight. Sepals ovate, 3–3.5 ´ 1–1.5 mm, margin membranous, lateral pair subsaccate; petals white, obovate, 7–10 ´ 3–4 mm, not clawed, apex rounded to emarginated; filaments slightly spreading, median pairs 4–5 mm, lateral pair 2.5–3.5 mm; anther narrowly oblong, 1–1.5 mm; ovules 14–18 per ovary. Fruit linear, strongly flattened, (1.5–)2–3(–3.5) cm ´ 1.5–2 mm; valves smooth, glabrous; styles slender, 1.5–3 mm. Seeds brown, oblong, 2–3 ´ 1–2 mm, winged all around; wing to 1 mm wide.

Flowering: Apr–Jun.

Habitat: moist places, streamsides.

Elevation: 0–1000 m.

Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hebei, Heilongjiang, Henan, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Nei Mongol, Shandong, Sichuan, Zhejiang), Japan, Korea, Russia (Far East, Siberia).