. Eudema nubigena Humboldt & Bonpland, Pl. Aequinoct. 2: 136. 1813.
Plants densely cespitose, cushion forming, with a stout, woody, much-branched caudex covered with petiolar remains of previous years. Leaves basal, densely rosulate, petiolate, numerous, somewhat fleshy, glabrous or ciliate with simple trichomes to 1.2 mm; petioles persistent, thick, (2-)3-10 mm; leaf blade spatulate to ovate or oblanceolate, (3-)4-8(-9) mm, 1.5-4(-5) mm wide, margin entire, apex obtuse to subacute. Sepals ovate to oblong, (2-)2.5-3.5(-6) mm, (1.2-)1.5-2 mm wide, erect, margin scarious; petals white to creamy white, broadly obovate, 3-5 mm, 1.5-2.5(-2.8) mm wide, not clawed; filaments erect, slender, 2-3(-3.5) mm; anthers usually violet, oblong, 0.7-1 mm. Fruiting pedicels glabrous, 2-5(-10) mm. Fruit broadly to narrowly ovoid, (2-)3-5(-6) mm, 2-3 mm wide, subterete, rounded at base and apex; valves glabrous, obscurely veined; septum reduced to a narrow rim; style slender, 0.4-2.5(-3) mm. Seeds (2-)4-8(-10) per fruit, dark brown, oblong-ovoid, (1.4-)1.6-2 mm, (0.8-)1-1.3(-1.4) mm wide, conspicuously reticulate; funicles thick, to 0.6 mm.
Two subspecies: Ecuador
Key to the subspecies of Eudema nubigena
1. Leaf blade conspicuously ciliate; fruit mostly narrowly ovoid; style 0.4-1.5(-3) mm .................4a. subsp. nubigena
1. Leaf blade not ciliate; fruit narrowly to broadly ovoid; style (1.2-)1.5-3 mm ............................4b. subsp. remyana